Cirsium pulcherrimum |
Cirsium helenioides |
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Wyoming thistle |
melancholy thistle |
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Habit | Perennials polycarpic, 15–60(–90) cm; deep-seated woody tap-roots and caudices. | Perennials, 40–120 cm; runner roots. | ||||
Stems | 1–few, erect or ascending, arachnoid-tomentose or ± glabrate; branches 0–5+, usually in distal 1/2, ascending. |
single, erect, ± arachnoid-tomentose; branches 0 or few, ascending. |
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Leaves | blades linear to oblong, oblanceolate, or elliptic, 5–25 × 0.6–7 cm, unlobed and merely spinulose or spiny-dentate to regularly pinnatifid, lobes 5–8(–many) pairs, well separated, usually with broad, U-shaped sinuses to crowded, linear to triangular-ovate, ascending-spreading to retrorse, merely spinulose to coarsely dentate or few lobed, main spines 2–7 mm, ± slender, abaxial faces gray to white, usually densely arachnoid-tomentose, sometimes ± glabrate, sometimes villous with septate trichomes along veins, adaxial green, glabrous or less commonly thinly to densely gray-tomentose; basal often present at flowering, spiny winged-petiolate; principal cauline well distributed, gradually reduced distally, proximal usually winged-petiolate, mid and distal sessile, bases decurrent as spiny wings 1.5–3.5 cm; distalmost reduced, ± bractlike. |
oblong to broadly lanceolate, 20–40 × 4–8 cm, finely spinulose-dentate or proximal cauline pinnatifid, lobes undivided, finely spinulose-dentate, main spines 1–2 mm, abaxial faces white-tomentose (with non-septate trichomes), adaxial glabrous; basal present at flowering, petiolate, bases tapered; cauline sessile, reduced distally, bases clasping, not decurrent; distal (few, well separated), oblong or linear, the uppermost reduced to linear bracts. |
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Peduncles | 0–15 cm. |
2–10(–30) cm (elevated above distal leaves). |
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Involucres | ovoid to campanulate, 1.8–2.7 × 1–2 cm, thinly arachnoid-tomentose or glabrate. |
broadly ovoid, 2–3 × 2–3.5 cm, glabrous or loosely arachnoid. |
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Corollas | pink to purple (creamy white), 18–25 mm, tubes 7–9 mm, throats 5.5–7.5 mm, lobes 4–8 mm; style tips 3–5.5 mm. |
red-purple, 25–30 mm, tubes 10–23 mm, throats 8–14 mm (noticeably wider than tubes), lobes 7–10 mm; style tips 4–5 mm. |
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Phyllaries | in 6–7 series, ± imbricate, green or with dark subapical patch or appendage, linear to linear-lanceolate, margins entire, abaxial faces with narrow glutinous ridge; outer and middle bases appressed, apical appendages spreading to stiffly ascending, linear-lanceolate to acicular, entire, spines spreading or ascending, stout, 2–7 mm, often flattened; apices of inner stiffly erect or sometimes flexuous, narrow, flat. |
in 8–10 series, imbricate, green, ovate or lanceolate (outer) to linear-lanceolate (inner), abaxial faces with a prominent elongate glutinous ridge, outer and middle tightly appressed, margins entire, apices with ascending, weak spines 0–1 mm; apices of inner phyllaries attenuate, flat. |
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Heads | 1–few, borne singly or in 2–3-headed clusters in ± congested flat-topped or racemiform arrays at tips of main stem and branches, sometimes also in distal axils. |
borne singly or less commonly 2–5 in terminal clusters. |
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Cypselae | tan to dark brown, 5–6 mm, apical collars yellow, narrow; pappi 14–16 mm. |
light brown, 3–5 mm, bodies and apical collars concolorous; pappi 20–30 mm. |
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2n | = 34. |
= 34. |
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Cirsium pulcherrimum |
Cirsium helenioides |
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Phenology | Flowering summer (Jul–Aug). | |||||
Habitat | Fjordlands | |||||
Elevation | 0–50 m (0–200 ft) | |||||
Distribution |
CO; ID; MT; NE; UT; WY
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Greenland; Iceland; Europe; Asia |
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Discussion | Varieties 2 (2 in the flora). Cirsium pulcherrimum is closely related to C. clavatum. In southeastern Wyoming and northern Colorado some plants combine foliage and involucral characters of C. pulcherrimum var. pulcherrimum and C. clavatum var. americanum. The inheritance of these characters needs to be examined at the population level to determine whether the intermediates are hybrids or the products of past introgression or incomplete differentiation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium helenioides is one of only two species of the genus that have native populations in the Old World and the flora area. Neither reaches the North American mainland. The conservation status of Cirsium helenioides is not known; it is known in the flora area only from a single fjord and possibly should be considered of conservation concern. of conservation concern (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 125. | FNA vol. 19, p. 110. | ||||
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Carduus pulcherrimus | Carduus helenioides, C. heterophyllum | ||||
Name authority | (Rydberg) K. Schumann: Just’s Bot. Jahresber. 29(1): 566. (1903) | (Linnaeus) Hill: Hort. Kew., 64. (1768) | ||||
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