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lost or Palo Alto thistle, lost thistle, Palo Alto thistle

Brewer's thistle, California swamp or Douglas' thistle, Douglas' thistle, swamp thistle

Habit Biennials or perennials, probably more than 100 cm; rootstock unknown. Biennials or short-lived monocarpic perennials, 60–250 cm; taprooted.
Stems

stout, erect, loosely arachnoid with fine trichomes and villous with jointed trichomes; branching unknown.

1–several, erect or ascending, densely gray-tomentose;

branches few–many, ascending to spreading.

Leaves

blades elliptic to oblanceolate, 15–30+ × 6–8+ cm, divided halfway or more to midveins, lobes linear-lanceolate, rigidly spreading, entire or trifid, acuminate, main spines stout, 5–15 mm, abaxial faces tomentose with fine, non-septate trichomes, villous along major veins with septate trichomes, adaxial glabrescent or sparsely tomentose, villous along veins;

basal not observed;

cauline well distributed, distally not much reduced, sessile, bases clasping, not decurrent.

blades oblong-elliptic to obovate, 10–60 × 2–15 cm, unlobed or shallowly to deeply pinnatifid, lobes lanceolate to ovate-triangular, ascending to spreading, entire to coarsely dentate or lobed, main spines slender to stout, 2–30 mm, faces densely gray-tomentose, rarely glabrate;

basal present at flowering, petiolate;

principal cauline well distributed, proximal winged-petiolate, distal sessile, bases auriculate-clasping or decurrent as a spiny wing 1–3 cm;

distalmost well separated, bractlike.

Peduncles

0–1 cm.

0–4(–8) cm.

Involucres

hemispheric to broadly campanulate, 3–4 × 4–5+ cm, arachnoid.

ovoid to hemispheric, 1.5–3 cm, 2–4.5 cm diam, loosely arachnoid on phyllary margins or glabrate.

Corollas

white, 30–33 mm, tubes 16 mm, throats 9–12 mm, lobes 5.5–9 mm;

style tips 6 mm.

rose-purple (white or pinkish-tinged), 18–21 mm, tubes 8–9 mm, throats 5–6 mm (abruptly expanded), lobes 5–6 mm;

style tips 3–4.5 mm.

Phyllaries

in 6–8 series, narrowly lanceolate to linear, outer subequal, rigidly spreading, spines 5–10 mm, inner ± imbricate, bodies appressed, glutinous ridge absent, apices spreading, margins spinulose or scabrid, apices of mid and inner flattened, spineless, scabrid.

in 6–8 series, imbricate, often with dark purple patch near tip, ovate-lanceolate (outer) to linear-lanceolate (inner), abaxial faces with linear to elliptic glutinous ridge;

outer and middle appressed, entire, spines spreading, 1–9 mm;

apices of inner often purple-tinged, often flexuous, flattened, spineless, scabrid.

Heads

1–5, terminal and in distal axils in spiciform arrays.

10–many, often crowded at branch tips, collectively forming paniculiform arrays.

Cypselae

light brown, 6 mm, collars also light brown, ca. 0.75 mm;

pappi 25–33 mm.

dark brown to black, 5–6 mm, apical collars not differentiated;

pappi 15–20 mm.

Cirsium praeteriens

Cirsium douglasii

Phenology Flowering summer (Jun–Jul).
Habitat Habitat unknown
Elevation 0–100 m (0–300 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Cirsium praeteriens is known only from Santa Clara County, where J. W. Congdon collected it in Palo Alto in 1897 and 1901. It is presumed extinct.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal cauline leaves with larger spines mostly less than 7 mm
var. breweri
1. Distal cauline leaves with larger spines usually 7–20 mm
var. douglasii
Source FNA vol. 19, p. 160. FNA vol. 19, p. 133.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. douglasii var. breweri, C. douglasii var. douglasii
Name authority J. F. Macbride: Contr. Gray Herb. 53: 19. (1918) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 643. (1838)
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