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lost or Palo Alto thistle, lost thistle, Palo Alto thistle

slough thistle

Habit Biennials or perennials, probably more than 100 cm; rootstock unknown. Annuals or biennials, (60–)100–300 cm; taprooted.
Stems

stout, erect, loosely arachnoid with fine trichomes and villous with jointed trichomes; branching unknown.

usually 1, erect, stout, (hollow, 2–10 cm diam. at base), openly branched distally, thinly arachnoid, villous with jointed trichomes, at least proximally.

Leaves

blades elliptic to oblanceolate, 15–30+ × 6–8+ cm, divided halfway or more to midveins, lobes linear-lanceolate, rigidly spreading, entire or trifid, acuminate, main spines stout, 5–15 mm, abaxial faces tomentose with fine, non-septate trichomes, villous along major veins with septate trichomes, adaxial glabrescent or sparsely tomentose, villous along veins;

basal not observed;

cauline well distributed, distally not much reduced, sessile, bases clasping, not decurrent.

blades elliptic to broadly oblanceolate, 15–70 × 30–150+ cm, flat, pinnatifid 1/2–2/3 distance to midvein, larger usually with broad sinuses, lobes broad, few lobed or dentate, main spines 3–8 mm, abaxial faces gray-tomentose, adaxial thinly arachnoid-tomentose, sometimes midveins with jointed trichomes;

basal present or withered at flowering, winged-petiolate;

principal cauline sessile, progressively reduced distally, bases clasping or short-decurrent 1–2 cm;

distal cauline reduced, becoming bractlike, sometimes spinier than proximal.

Peduncles

0–1 cm.

0–15 cm.

Involucres

hemispheric to broadly campanulate, 3–4 × 4–5+ cm, arachnoid.

ovoid to campanulate, 1.5–3 × 1.5–3 cm, ± glabrous.

Corollas

white, 30–33 mm, tubes 16 mm, throats 9–12 mm, lobes 5.5–9 mm;

style tips 6 mm.

pale rose-purple (white), 19–26 mm, tubes 9–12 mm, throats 4–6 mm, lobes 5–9 mm;

style tips 3.5–4.5 mm.

Phyllaries

in 6–8 series, narrowly lanceolate to linear, outer subequal, rigidly spreading, spines 5–10 mm, inner ± imbricate, bodies appressed, glutinous ridge absent, apices spreading, margins spinulose or scabrid, apices of mid and inner flattened, spineless, scabrid.

in 5–7 series, weakly unequal, dark green to brownish, lanceolate (outer) to linear (inner), abaxial faces without (or with very obscure) glutinous ridge;

outer and middle appressed or apices spreading, at least outer irregularly spiny-fringed, finely serrulate, spines slender, 3–5 mm;

apices of inner erect, abaxial faces gray-tomentose, ± twisted.

Heads

1–5, terminal and in distal axils in spiciform arrays.

1–several at branch tips, closely subtended by clustered leafy bracts or not, collectively forming open, corymbiform or paniculiform arrays.

Cypselae

light brown, 6 mm, collars also light brown, ca. 0.75 mm;

pappi 25–33 mm.

dark brown, 5–5.5 mm, collars narrow, ± stramineous;

pappi 15–20 mm.

2n

= 32.

Cirsium praeteriens

Cirsium crassicaule

Phenology Flowering summer (Jun–Jul). Flowering spring (Apr–Jun).
Habitat Habitat unknown Freshwater marshes, canal banks
Elevation 0–100 m (0–300 ft) 5–100 m (0–300 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Of conservation concern.

Cirsium praeteriens is known only from Santa Clara County, where J. W. Congdon collected it in Palo Alto in 1897 and 1901. It is presumed extinct.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cirsium crassicaule is known only from a few sites in the San Joaquin Valley. Some populations are threatened by habitat modification and development pressures.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 160. FNA vol. 19, p. 132.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Synonyms Carduus crassicaulis
Name authority J. F. Macbride: Contr. Gray Herb. 53: 19. (1918) (Greene) Jepson: Fl. W. Calif., 506. (1901)
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