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dune thistle, Pitcher's thistle, sand-dune thistle

pasture thistle

Habit Biennials or short-lived monocarpic perennials, 20–100 cm; taproots long. Biennials or monocarpic perennials, 25–100 cm; sometimes perennating by root sprouts.
Stems

1 or few, erect, densely gray-tomentose;

branches 0 to several, ascending to spreading.

erect, villous with septate trichomes and sometimes thinly arachnoid tomentose;

branches 0–few, distal, ascending.

Leaves

blades elliptic to obovate, 10–30 × 8–14 cm, deeply divided nearly to midveins, lobes ascending to spreading, linear, remote, margins revolute, entire or minutely spinulose, main spines 1–2 mm, faces gray-tomentose, more densely so abaxially;

basal present or withered at flowering, petiolate;

principal cauline well distributed, bases decurrent as linear-lobed to spiny wings 1–3 cm;

distal cauline well developed.

blades oblong-elliptic, 5–30+ × 2–10 cm, ± undulate, shallowly to deeply pinnatifid, lobes ovate to broadly triangular, usually separated by broad sinuses, spinose-dentate or lobed, main spines 1.5–7 mm, slender to stout, abaxial faces villous with septate trichomes, at least along veins, sometimes thinly arachnoid, adaxial faces villous with septate trichomes and thinly arachnoid tomentose or subglabrous;

basal often present at flowering, bases tapered;

principal cauline sessile, moderately reduced distally, bases often auriculate-clasping;

distal reduced, similar to proximal.

Peduncles

0–5 cm.

0–15+ cm (above distal leaves), leafy-bracted.

Involucres

ovoid to campanulate, 2–3 × 2–3 cm, loosely arachnoid on phyllary margins or glabrate.

broadly cylindric to ovoid, 3.5–5 cm, 2.5–3 cm diam. (appearing much wider and hemispheric to campanulate in pressed specimens), loosely arachnoid on phyllary margins or glabrate.

Corollas

dull white or pinkish-tinged (rarely rich purple), 20–30 mm, tubes 8.5–15 mm, throats 4.5–10 mm, lobes 3–8 mm;

style tips 3.5–5.5 mm.

pink to purple (white), 40–60 mm, tubes 20–35 mm, throats 11–15 mm, lobes 7–10 mm;

style tips 3–7 mm.

Phyllaries

in 6–8 series, imbricate, ovate-lanceolate (outer) to linear-lanceolate (inner), abaxial faces with narrow glutinous ridge;

outer and middle appressed, acute, spines ascending to spreading, slender, 1–2(–3) mm;

apices of inner often flexuous, flattened, spineless, scabrid.

in 8–10 series, imbricate, ovate or lanceolate (outer) to lance-linear (inner), abaxial faces with ± prominent glutinous ridge, outer and middle appressed, apices ascending to spreading, spines 1.5–6 mm;

apices of middle and inner narrowed and scabrid-denticulate, innermost spineless, tapered and entire or with expanded, erose-denticulate, flexuous tips.

Heads

1–20+ in corymbiform arrays.

1–few, borne singly at tips of main stem and branches, often closely subtended by 1–several bracts.

Cypselae

light brown, sometimes with darker streaks, 6–7.5 mm, apical collars lighter colored, very narrow;

pappi 15–30 mm, usually noticeably shorter than corolla.

stramineous to brown, 3.5–5 mm, apical collars yellow or colored like body;

pappi 35–45 mm.

2n

= 34.

= 30.

Cirsium pitcheri

Cirsium pumilum

Phenology Flowering spring–summer (May–Sep).
Habitat Sand dunes and beaches
Elevation 180–200 m (600–700 ft)
Distribution
from FNA
IL; IN; MI; WI; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; DE; IA; IL; IN; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; SC; VA; VT; WI; WV; ON; United States
[WildflowerSearch map]
[BONAP county map]
Discussion

Cirsium pitcheri is endemic to beach and dune habitats around lakes Huron, Michigan, and Superior. It has been extirpated from portions of its former range at the southern end of Lake Michigan. It is threatened by foot traffic, off-road vehicular activity, and clearing and development of beachside habitats. It is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Flowers of Cirsium pumilum are reportedly sweetly scented.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spines of outer phyllaries stout, 3–6 mm; plants usually with several long branches; leaves usually lobed nearly to the midvein; roots rarely and only slightly tuberous-thickened; cypselae usually 3.5– 4 mm
var. pumilum
1. Spines of outer phyllaries slender, 1.5–3 mm; plants simple or distally with a few short branches; leaves usually shallowly lobed; roots often tuberous-thickened; cypselae usually 4.5–5 mm
var. hillii
Source FNA vol. 19, p. 122. FNA vol. 19, p. 116.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. pumilum var. hillii, C. pumilum var. pumilum
Synonyms Cnicus pitcheri Carduus pumilus
Name authority (Torrey ex Eaton) Torrey & A. Gray: Fl. N. Amer. 2: 456. (1843) (Nuttall) Sprengel: Syst. Veg. 3: 375. (1826)
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