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dune thistle, Pitcher's thistle, sand-dune thistle

bristly or horrid or yellow or bull thistle, bristly thistle, bull thistle, horrid thistle, yellow thistle

Habit Biennials or short-lived monocarpic perennials, 20–100 cm; taproots long. Biennials or perennials, (± fleshy), 15–250 cm; stout taproots and a fascicle of fleshy lateral roots, often perennating by root sprouts.
Stems

1 or few, erect, densely gray-tomentose;

branches 0 to several, ascending to spreading.

1–several, usually erect, often stout, glabrous to densely tomentose;

branches 0–many, spreading to ascending, short, stout.

Leaves

blades elliptic to obovate, 10–30 × 8–14 cm, deeply divided nearly to midveins, lobes ascending to spreading, linear, remote, margins revolute, entire or minutely spinulose, main spines 1–2 mm, faces gray-tomentose, more densely so abaxially;

basal present or withered at flowering, petiolate;

principal cauline well distributed, bases decurrent as linear-lobed to spiny wings 1–3 cm;

distal cauline well developed.

blades linear to oblanceolate or oblong-elliptic, 10–40 × 2–10 cm, unlobed and spiny-dentate to deeply pinnatifid, lobes spiny-dentate or coarsely lobed, main spines stout, 5–30 mm, abaxial faces subglabrous to loosely tomentose, adaxial glabrous to ± densely villous with septate trichomes;

basal present at flowering, spiny winged-petiolate, bases often tapered;

principal cauline sessile, well distributed, often not much reduced distally, bases often ± auriculate-clasping;

distal cauline often spinier than the proximal.

Peduncles

0–5 cm.

0–5 cm.

Involucres

ovoid to campanulate, 2–3 × 2–3 cm, loosely arachnoid on phyllary margins or glabrate.

hemispheric to campanulate, 3–5 × 3–8 cm.

Corollas

dull white or pinkish-tinged (rarely rich purple), 20–30 mm, tubes 8.5–15 mm, throats 4.5–10 mm, lobes 3–8 mm;

style tips 3.5–5.5 mm.

white to yellow, pink, purple, or red, 30–47 mm, tubes 11–30 mm, throats 6–10 mm, lobes 7–10 mm;

style tips 3–5 mm.

Phyllaries

in 6–8 series, imbricate, ovate-lanceolate (outer) to linear-lanceolate (inner), abaxial faces with narrow glutinous ridge;

outer and middle appressed, acute, spines ascending to spreading, slender, 1–2(–3) mm;

apices of inner often flexuous, flattened, spineless, scabrid.

in 5–9 series, subequal to imbricate, light green to stramineous, lanceolate to linear, distally often with reddish margins, abaxial faces without glutinous ridge, often ± thinly tomentose, often scabridulous in submarginal bands;

outer and middle appressed-ascending, bodies usually reddish-tinged, margins setulose-ciliolate, apices acuminate, spines 1–2 mm, weak;

apices of inner straight, flat.

Heads

1–20+ in corymbiform arrays.

1–20 in subcapitate to corymbiform arrays (each closely subtended by an involucre-like ring of spiny-margined bracts).

Cypselae

light brown, sometimes with darker streaks, 6–7.5 mm, apical collars lighter colored, very narrow;

pappi 15–30 mm, usually noticeably shorter than corolla.

straw-colored to tan, 4–6 mm, apical collars weakly differentiated;

pappi 25–35 mm.

2n

= 34.

= 32, 33, 34, 35.

Cirsium pitcheri

Cirsium horridulum

Phenology Flowering spring–summer (May–Sep).
Habitat Sand dunes and beaches
Elevation 180–200 m (600–700 ft)
Distribution
from FNA
IL; IN; MI; WI; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; FL; GA; LA; MA; MD; ME; MS; NC; NH; NJ; NY; OK; PA; RI; SC; TN; TX; VA; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Cirsium pitcheri is endemic to beach and dune habitats around lakes Huron, Michigan, and Superior. It has been extirpated from portions of its former range at the southern end of Lake Michigan. It is threatened by foot traffic, off-road vehicular activity, and clearing and development of beachside habitats. It is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (3 in the flora).

Although several variants have been given taxonomic recognition as species, these seem at most races. Flower color varies greatly, sometimes within populations and sometimes on a populational or regional basis. Herbarium specimens are sometimes difficult to assign to variety.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems densely tomentose; involucres ± densely tomentose
var. horridulum
1. Stems glabrous or sparsely tomentose; involucres glabrous
→ 2
2. Leaves shallowly to deeply pinnatifid, main spines 10–30 mm
var. megacanthum
2. Leaves spinose-dentate to shallowly pinnatifid, main spines mostly 5–10 mm
var. vittatum
Source FNA vol. 19, p. 122. FNA vol. 19, p. 114.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. horridulum var. horridulum, C. horridulum var. megacanthum, C. horridulum var. vittatum
Synonyms Cnicus pitcheri
Name authority (Torrey ex Eaton) Torrey & A. Gray: Fl. N. Amer. 2: 456. (1843) Michaux: Fl. Bor.-Amer. 2: 90. (1803)
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