Cirsium palustre |
Cirsium undulatum |
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cirse ou chardon des marais, European Marsh thistle, European swamp or marsh thistle, European swamp thistle, marsh thistle |
gray thistle, wavy-leaf thistle, wavyleaf or gray or pasture thistle |
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Habit | Biennials or monocarpic perennials, 30–200(–300) cm; clusters of fibrous roots. | Perennials, 20–230 cm; deeply seated runner roots that produce adventitious buds. |
Stems | single, erect, villous to tomentose with jointed trichomes, distally tomentose with fine, unbranched trichomes; branches 0–few, ascending, (short). |
1–several, erect or ascending, densely gray-tomentose; branches 0–few, usually above middle, ascending. |
Leaves | blades narrowly elliptic to oblanceolate, 15–30+ × 3–10 cm, margins shallowly to very deeply pinnatifid, narrow lobes separated by broad sinuses, spiny-dentate to lobed, main spines 2–6 mm, abaxial villous to tomentose with jointed trichomes, sometimes also thinly tomentose with fine unbranched trichomes, adaxial faces villous with septate trichomes or glabrate; basal often present at flowering, petioles spiny-winged, bases tapered; cauline many, sessile, gradually reduced and becoming widely spaced above, bases long-decurrent with prominently spiny wings; distal cauline deeply pinnatifid with few-toothed spine-tipped lobes. |
blades elliptic to oblong or ovate, 10–40 × 1–10 cm, margins strongly undulate, coarsely dentate or shallowly to deeply lobed, lobes ascending to spreading, ± triangular, well separated to closely spaced, spinulose and coarsely dentate or usually cleft into 2–3 lanceolate to triangular, often entire-margined, spine-tipped divisions, main spines (yellowish), 2–12+ mm, abaxial densely gray-tomentose, adaxial faces thinly tomentose; basal sometimes present at flowering, winged-petiolate; principal cauline becoming sessile and progressively reduced distally, widest at base, bases ± auriculate-clasping to short-decurrent; distal reduced, spinier. |
Peduncles | 0–1 cm. |
0–25+ cm. |
Involucres | ovoid to campanulate, 1–1.5 × 0.8–1.3 cm, thinly cobwebby tomentose with fine unbranched trichomes. |
ovoid to hemispheric or broadly campanulate, 2.5–4.5 × 1.5–4.5 cm, loosely arachnoid on phyllary margins or glabrate. |
Corollas | lavender to purple (white), 11–13 mm, tubes 5–7 mm, throats 2–3 mm, lobes 3–4.5 mm; style tips 1.5–2 mm. |
lavender to pink, purple, or white, 24–50 mm, tubes 12–28 mm, throats 6–14 mm, lobes 6.5–13 mm; style tips 5–7.5 mm. |
Phyllaries | in 5–7 series, strongly imbricate, greenish, or with purplish tinge, lanceolate to ovate (outer) or linear-lanceolate (inner), margins thinly arachnoid-ciliate, abaxial faces with narrow glutinous ridge, outer and middle appressed, entire, apices acute, mucronate or spines erect or spreading, weak, 0.3–1 mm; apices of inner phyllaries purplish, linear-attenuate, scarious, flat. |
in 8–12 series, imbricate, ovate to lanceolate (outer) to linear-lanceolate (inner), abaxial faces with prominent glutinous ridge; outer and middle appressed, spines spreading, 1.5–5 mm; apices of inner narrow, often flexuous, flat, ± entire, spineless or weakly spiny. |
Heads | few–many in dense clusters at branch tips. |
1–10+, terminal on branches, in leafy, ± corymbiform arrays. |
Cypselae | tan to stramineous, 2.5–3.5 mm, apical collars 0.1–0.2 mm, shiny; pappi 9–11 mm. |
light to dark brown, 6–7 mm, bodies and apical collars concolorous, narrow; pappi 20–38 mm (usually scabridulous). |
2n | = 34. |
= 26. |
Cirsium palustre |
Cirsium undulatum |
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Phenology | Flowering summer (Jul–Aug). | Flowering spring–autumn (May–Oct). |
Habitat | Marshes, wet forests | Mixedgrass prairie, shortgrass prairie, Palouse prairie, sagebrush deserts, pinyon-juniper woodlands, openings in montane coniferous forests, often in disturbed areas |
Elevation | 10–800 m (0–2600 ft) | 100–2800 m (300–9200 ft) |
Distribution |
MA; MI; NH; NY; WI; BC; NF; NS; ON; QC; SPM; Europe [Introduced in North America]
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AZ; CA; CO; GA; IA; ID; IL; IN; KS; MI; MN; MO; MT; ND; NE; NM; OK; OR; PA; SD; TX; UT; WA; WI; WY; AB; BC; MB; SK; Mexico (Chihuahua, Coahuila, Durango, Sonora)
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Discussion | Cirsium palustre is a noxious weed, native to Europe, that invasively spreads through wetland communities, forming impenetrable spiny stands as it displaces native species. The range of this pernicious weed in North America is rapidly expanding. It has the potential to spread into boreal forest areas across the continent; in Europe it grows nearly to the Arctic Circle. The rapid spread of C. palustre in Michigan (E. G. Voss 1972–1996, vol. 3) is indicative of its invasiveness. Spontaneous hybrids between C. palustre and C. arvense have been reported from England and other European countries (W. A. Sledge 1975) and can be expected wherever these species grow together in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium undulatum is widely distributed in the wstern half of North America from the dry plains and plateaus of the Pacific Northwest eastward across the Great Plains to Manitoba and the Dakotas and south to Texas, New Mexico, and northwestern Mexico. It occurs in scattered localities in the Rocky Mountains and northeastern Great Basin region. At least some of the few widely scattered records from the eastern United States are probably introductions. Cirsium undulatum is both widespread and variable. Plants of the Great Plains region tend to be low-growing with a few large heads and elongate corollas. Plants of the Pacific Northwest are usually taller and produce smaller, more numerous heads with shorter corollas. A detailed study of this species might reveal races worthy of recognition as infraspecific taxa. Wavyleaf thistle is listed by California as a noxious weed. However, most reports of Cirsium undulatum in California are based upon misidentifications of C. canescens. Cirsium undulatum is known to hybridize with C. flodmanii, C. hookerianum, and C. scariosum var. coloradense. J. T. Howell (1960b) reported that C. undulatum was suspected to hybridize with C. brevifolium in the Pacific Northwest. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 110. | FNA vol. 19, p. 120. |
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium |
Sibling taxa | ||
Synonyms | Carduus palustris | Carduus undulatus, C. megacephalum, C. undulatum var. megacephalum |
Name authority | (Linnaeus) Scopoli: Fl. Carniol. ed. 2, 2: 128. (1772) | (Nuttall) Sprengel: Syst. Veg. 3: 374. (1826) |
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