Cirsium palustre |
Cirsium longistylum |
|
---|---|---|
cirse ou chardon des marais, European Marsh thistle, European swamp or marsh thistle, European swamp thistle, marsh thistle |
long-style thistle |
|
Habit | Biennials or monocarpic perennials, 30–200(–300) cm; clusters of fibrous roots. | Perennials monocarpic, 40–150 cm; taprooted. |
Stems | single, erect, villous to tomentose with jointed trichomes, distally tomentose with fine, unbranched trichomes; branches 0–few, ascending, (short). |
usually 1, erect, less commonly several, ascending, simple to sparingly short-branched in distal 1/2, less commonly openly branched, villous with jointed trichomes; branches on distal stems 0–many, short, ascending. |
Leaves | blades narrowly elliptic to oblanceolate, 15–30+ × 3–10 cm, margins shallowly to very deeply pinnatifid, narrow lobes separated by broad sinuses, spiny-dentate to lobed, main spines 2–6 mm, abaxial villous to tomentose with jointed trichomes, sometimes also thinly tomentose with fine unbranched trichomes, adaxial faces villous with septate trichomes or glabrate; basal often present at flowering, petioles spiny-winged, bases tapered; cauline many, sessile, gradually reduced and becoming widely spaced above, bases long-decurrent with prominently spiny wings; distal cauline deeply pinnatifid with few-toothed spine-tipped lobes. |
blades linear to oblong or elliptic, 10–30+ × 1–10 cm, margins flat to undulate, subentire to coarsely dentate or shallowly to deeply pinnatifid, lobes lance-oblong to broadly triangular, spinulose to spiny-dentate or shallowly lobed, main spines 3–12 mm, abaxial faces green and subglabrous to gray- or white-tomentose with felted arachnoid trichomes, ± villous to tomentose along major veins with septate trichomes, rarely glabrous or glabrate, adaxial ± green, glabrous or villous with septate trichomes; basal often present at flowering, spiny winged-petiolate or sessile; principal cauline well distributed, proximally winged-petiolate, distally sessile, gradually reduced, bases sometimes short-decurrent (0–2 cm); distal ± reduced, often narrower than the proximal, sometimes with non-pigmented bases. |
Peduncles | 0–1 cm. |
0–15+ cm. |
Involucres | ovoid to campanulate, 1–1.5 × 0.8–1.3 cm, thinly cobwebby tomentose with fine unbranched trichomes. |
(green), broadly ovoid, 1.5–2.5 × 1.5–2.5 cm, loosely villous with septate trichomes, sparingly if at all arachnoid. |
Corollas | lavender to purple (white), 11–13 mm, tubes 5–7 mm, throats 2–3 mm, lobes 3–4.5 mm; style tips 1.5–2 mm. |
white, ochroleucous, 19–23 mm, tubes 6.5–8.5 mm, throats 7.5–11 mm, lobes 4–5 mm; style tips 4–5.5 mm, conspicuously exserted. |
Phyllaries | in 5–7 series, strongly imbricate, greenish, or with purplish tinge, lanceolate to ovate (outer) or linear-lanceolate (inner), margins thinly arachnoid-ciliate, abaxial faces with narrow glutinous ridge, outer and middle appressed, entire, apices acute, mucronate or spines erect or spreading, weak, 0.3–1 mm; apices of inner phyllaries purplish, linear-attenuate, scarious, flat. |
in 4–8 series, subequal, ± lanceolate, bases appressed, apices ascending, linear to broadly expanded, erose to lacerate or spiny-fringed, spines straight, slender, 2–3 mm, abaxial faces with or without narrow glutinous ridge; apices of inner flexuous, sometimes expanded and erose. |
Heads | few–many in dense clusters at branch tips. |
several–many, erect, usually in racemiform or spiciform arrays, usually closely subtended by clustered ± leafy bracts. |
Cypselae | tan to stramineous, 2.5–3.5 mm, apical collars 0.1–0.2 mm, shiny; pappi 9–11 mm. |
brown, 5.5–6.5 mm, apical collars not differentiated; pappi 17–20 mm. |
2n | = 34. |
= 34. |
Cirsium palustre |
Cirsium longistylum |
|
Phenology | Flowering summer (Jul–Aug). | Flowering summer (Jun–Aug). |
Habitat | Marshes, wet forests | Moist soil, roadsides, meadows, forest edges and openings |
Elevation | 10–800 m (0–2600 ft) | 1500–2400 m (4900–7900 ft) |
Distribution |
MA; MI; NH; NY; WI; BC; NF; NS; ON; QC; SPM; Europe [Introduced in North America]
|
MT |
Discussion | Cirsium palustre is a noxious weed, native to Europe, that invasively spreads through wetland communities, forming impenetrable spiny stands as it displaces native species. The range of this pernicious weed in North America is rapidly expanding. It has the potential to spread into boreal forest areas across the continent; in Europe it grows nearly to the Arctic Circle. The rapid spread of C. palustre in Michigan (E. G. Voss 1972–1996, vol. 3) is indicative of its invasiveness. Spontaneous hybrids between C. palustre and C. arvense have been reported from England and other European countries (W. A. Sledge 1975) and can be expected wherever these species grow together in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Cirsium longistylum is endemic to the Big Belt, Castle, Elkhorn, and Little Belt ranges of west-central Montana. It is highly variable, and several authors have suggested that it has introgressed with one or more other species (R. J. Moore and C. Frankton 1963; J. M. Poole and B. L. Heidel 1993; S. J. Brunsfeld and C. T. Baldwin, unpubl.). It is closely related to C. hookerianum, and the two probably share a common ancestry or a history of hybrid interactions dating back to the Pleistocene. Cirsium longistylum is perhaps also affected by modern or historic introgression involving C. scariosum var. scariosum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 110. | FNA vol. 19, p. 149. |
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium |
Sibling taxa | ||
Synonyms | Carduus palustris | |
Name authority | (Linnaeus) Scopoli: Fl. Carniol. ed. 2, 2: 128. (1772) | R. J. Moore & Frankton: Canad. J. Bot. 41: 1562, plate 1. (1963) |
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