Flora of North America species comparison

single, erect, villous to tomentose with jointed trichomes, distally tomentose with fine, unbranched trichomes;

branches 0–few, ascending, (short).

usually single, erect, glabrous to ± tomentose, sometimes sparsely villous with septate trichomes;

branches few, usually distal, ascending.

blades narrowly elliptic to oblanceolate, 15–30+ × 3–10 cm, margins shallowly to very deeply pinnatifid, narrow lobes separated by broad sinuses, spiny-dentate to lobed, main spines 2–6 mm, abaxial villous to tomentose with jointed trichomes, sometimes also thinly tomentose with fine unbranched trichomes, adaxial faces villous with septate trichomes or glabrate;

basal often present at flowering, petioles spiny-winged, bases tapered;

cauline many, sessile, gradually reduced and becoming widely spaced above, bases long-decurrent with prominently spiny wings;

distal cauline deeply pinnatifid with few-toothed spine-tipped lobes.

blades linear to oblanceolate or elliptic, 10–30 × 1–5 cm, unlobed and spinulose to irregularly dentate or pinnatifid, lobes narrowly to broadly triangular, sometimes coarsely toothed or lobed toward base, acuminate, main spines slender, 1–5 mm, abaxial faces gray-tomentose, adaxial glabrous or sparsely villous with septate trichomes;

basal often present at flowering, petioles slender, winged, bases long-tapered;

principal cauline relatively few (10–25), petiolate or distal sessile, mostly restricted to proximal 1/2 of stems, progressively reduced distally, bases tapered, not decurrent;

distal cauline widely separated, linear to narrowly elliptic, reduced, becoming ± bractlike, merely spinulose to irregularly dentate or shallowly lobed.

0–1 cm.

slender, 1–15 cm (not overtopped by distal leaves).

ovoid to campanulate, 1–1.5 × 0.8–1.3 cm, thinly cobwebby tomentose with fine unbranched trichomes.

narrowly ovoid to campanulate, 1.2–2 × 1.2–2 cm, thinly arachnoid-ciliate.

lavender to purple (white), 11–13 mm, tubes 5–7 mm, throats 2–3 mm, lobes 3–4.5 mm;

style tips 1.5–2 mm.

pink-purple (white), 15–20 mm, tubes 5–9 mm, throats 5–7 mm (noticeably wider than tubes), lobes 4–5 mm;

style tips 4 mm.

in 5–7 series, strongly imbricate, greenish, or with purplish tinge, lanceolate to ovate (outer) or linear-lanceolate (inner), margins thinly arachnoid-ciliate, abaxial faces with narrow glutinous ridge, outer and middle appressed, entire, apices acute, mucronate or spines erect or spreading, weak, 0.3–1 mm;

apices of inner phyllaries purplish, linear-attenuate, scarious, flat.

in 7–10 series, imbricate, green, linear to lanceolate (outer) or linear to linear-lanceolate (inner), abaxial faces with narrow, glutinous ridge;

outer and middle ascending to appressed, bodies entire, apices widely spreading (at least the outer), spines ascending to spreading (at least the outer), slender, 1–4 mm;

apices of inner phyllaries flat, often twisted, acuminate.

few–many in dense clusters at branch tips.

(1–)2–9(–many), in paniculiform arrays.

tan to stramineous, 2.5–3.5 mm, apical collars 0.1–0.2 mm, shiny;

pappi 9–11 mm.

light brown, 3–4 mm, apical collars yellowish, 0.5–1 mm;

pappi 12–14 mm.

= 34.

= 20, 22.