Flora of North America species comparison

single, erect, villous to tomentose with jointed trichomes, distally tomentose with fine, unbranched trichomes;

branches 0–few, ascending, (short).

1–several, erect, thinly gray-tomentose with fine, non-septate trichomes;

branches 0–many, ascending.

blades narrowly elliptic to oblanceolate, 15–30+ × 3–10 cm, margins shallowly to very deeply pinnatifid, narrow lobes separated by broad sinuses, spiny-dentate to lobed, main spines 2–6 mm, abaxial villous to tomentose with jointed trichomes, sometimes also thinly tomentose with fine unbranched trichomes, adaxial faces villous with septate trichomes or glabrate;

basal often present at flowering, petioles spiny-winged, bases tapered;

cauline many, sessile, gradually reduced and becoming widely spaced above, bases long-decurrent with prominently spiny wings;

distal cauline deeply pinnatifid with few-toothed spine-tipped lobes.

oblanceolate or elliptic, 15–45 × 2–10 cm, unlobed and merely spinulose to dentate or deeply pinnatifid, lobes well separated, linear to triangular-ovate, merely spinulose to few toothed or lobed near base, margins often revolute, main spines 2–3(–6) mm, abaxial faces densely gray-tomentose, adaxial green, thinly tomentose or ± glabrate;

basal often present at flowering, narrowly winged-petiolate;

principal cauline well distributed, gradually reduced distally, bases of proximal cauline winged-petiolate or sessile, bases of distal cauline expanded and ± clasping, margins sometimes spinier than those of proximal;

distalmost cauline becoming bractlike, often unlobed or less deeply divided than proximal.

0–1 cm.

1–8 cm.

ovoid to campanulate, 1–1.5 × 0.8–1.3 cm, thinly cobwebby tomentose with fine unbranched trichomes.

hemispheric to campanulate, 2.5–3.5 × 2–4 cm, glabrous or loosely floccose.

lavender to purple (white), 11–13 mm, tubes 5–7 mm, throats 2–3 mm, lobes 3–4.5 mm;

style tips 1.5–2 mm.

creamy white, rarely lavender-tinged, 22–28 mm, tubes 8–13 mm, throats 7–11 mm, lobes 4–6 mm;

style tips 5–6 mm.

in 5–7 series, strongly imbricate, greenish, or with purplish tinge, lanceolate to ovate (outer) or linear-lanceolate (inner), margins thinly arachnoid-ciliate, abaxial faces with narrow glutinous ridge, outer and middle appressed, entire, apices acute, mucronate or spines erect or spreading, weak, 0.3–1 mm;

apices of inner phyllaries purplish, linear-attenuate, scarious, flat.

in 6–10 series, strongly imbricate, greenish to brown, ovate to lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge;

outer and middle appressed, bodies entire, spines abruptly spreading, fine, 2–3(–5) mm;

apices of inner commonly flexuous or reflexed, flat, scarious.

few–many in dense clusters at branch tips.

borne singly and terminal on main stems and branches or few from distal axils in corymbiform or paniculiform arrays.

tan to stramineous, 2.5–3.5 mm, apical collars 0.1–0.2 mm, shiny;

pappi 9–11 mm.

brown, 5–6 mm, apical collars yellowish, 0.5–1 mm;

pappi 18–22 mm.

= 34.

= 22, 26.