Cirsium occidentale var. californicum |
Cirsium occidentale var. venustum |
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California thistle, cobwebby thistle |
cobwebby thistle, Venus thistle |
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Habit | Plants erect, usually 50–200 cm, thinly to densely gray-tomentose, sometimes glabrate. | Plants usually erect, usually 50–300 cm, variably tomentose, sometimes ± glabrate. |
Leaf | faces abaxially green to gray, adaxially gray. |
|
Involucres | usually about as wide as long, 1.5–5 cm, subglabrous to densely arachnoid. |
usually longer than wide, 2–6 cm, subglabrous to densely arachnoid, usually without fine trichomes connecting tips of adjacent phyllaries. |
Corollas | white to light purple or rose, 18–35 mm. |
usually ± red (white, pink, rarely purple), 23–35 mm. |
Phyllaries | usually imbricate, mid apices appressed to loosely spreading or ascending, sometimes twisted, usually less than 1 cm (but sometimes much longer), 1–3 mm. |
imbricate, outer and mid apices ascending to rigidly spreading or reflexed, straight, 5–20+ × usually 2–3 mm. |
Heads | in ± open clusters, short- to long-pedunculate, elevated well above proximal leaves. |
sometimes in tight clusters at ends of peduncles, usually long-pedunculate, elevated well above proximal leaves. |
2n | = 28, 29, 30 (as C. californicum). |
= 30. |
Cirsium occidentale var. californicum |
Cirsium occidentale var. venustum |
|
Phenology | Flowering spring–summer (Apr–Aug). | Flowering spring–summer (Apr–Jul). |
Habitat | Pine-oak woodlands, riparian woodlands, chaparral, openings in mixed evergreen forests, roadsides | Foothill oak-pine woodlands, grasslands, chaparral, pinyon-juniper woodlands, Joshua tree woodlands, roadsides |
Elevation | 100–2200 m (300–7200 ft) | 200–2300 m (700–7500 ft) |
Distribution |
CA |
CA; NV |
Discussion | Variety californicum occurs in both coastal and interior mountains of California from the northern South Coast Range and the northern Sierra Nevada to the mountains of southwestern California. Considerable variation exists in head size, corolla color, and in length and display of phyllary appendages. In several areas of its range, the predominantly white- to light purple-flowered var. californicum occurs with red-flowered var. venustum. These plants are highly interfertile (H. Wells 1983; D. J. Keil and C. E. Turner 1992). Introgressive hybridization among them has resulted in a variety of emergent phenotypes and may have contributed to the variation within var. californicum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Variety venustum has the widest ecological range of the races of C. occidentale. Populations occur within a few miles of the California coast in the North and South Coast Ranges and western Transverse Range and range eastward across the state into scattered sites in the Sierra Nevada to the higher elevations of the arid mountains of the western Mojave Desert and adjacent areas of the southwestern Great Basin Desert. Most populations of these plants can be recognized by their striking red to reddish pink corollas. The heads are sometimes visited by hummingbirds as well as by a variety of insects. Intermediates have been documented between var. venustum and vars. californicum and candidissimum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 139. | FNA vol. 19, p. 140. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. californicum, C. bernardinum, C. californicum var. bernardinum, C. californicum subsp. pseudoreglense | Carduus venustus, C. occidentale subsp. venustum, C. proteanum |
Name authority | (A. Gray) D. J. Keil & C. E. Turner: Phytologia 73: 315. (1992) | (Greene) Jepson: Man. Fl. Pl. Calif., 1167. (1925) |
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