Cirsium nuttallii |
Cirsium discolor |
|
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Nuttall's thistle |
chardon discolore, field thistle |
|
Habit | Biennials, 20–350 cm; taprooted. | Biennials or sometimes perennials, 80–200 cm; taproots and often cluster of coarse fibrous roots, roots without tuberlike enlargements. |
Stems | usually single, erect, glabrous or villous with septate trichomes; branches few–many, ascending. |
single, erect, villous with septate trichomes, sometimes ± glabrate, distally ± tomentose; branches few–many, ascending. |
Leaves | blades narrowly to broadly elliptic, (10–)15–60 × (2–)5–15 cm, thin, ± flexible, deeply pinnatifid, lobes narrow, spreading, coarsely dentate or lobed, main spines 2–5 mm, abaxial faces thinly tomentose but often wholly glabrate in age, adaxial glabrous or sparsely villous with septate trichomes; basal often absent at flowering, petioles slender, winged, bases tapered; principal cauline becoming sessile and gradually reduced distally, bases spiny-lobed, sometimes decurrent; distal reduced to linear bracts. |
blades oblanceolate to elliptic or ovate, 10–25(–50) × 1–13 (–25) cm, usually deeply divided more than halfway to midveins, proximal sometimes undivided, lobes linear-lanceolate, margins revolute, ascending, entire or spinulose to remotely few toothed or sharply lobed, main spines 1–5 mm, abaxial faces white-tomentose, adaxial faces green, villous with septate trichomes or glabrate; basal usually absent at flowering, winged-petiolate, bases tapered; principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping, not decurrent; distal cauline well developed. |
Peduncles | 1–15 cm, essentially naked (not overtopped by crowded distal leaves). |
0–5 cm (not overtopped by crowded distal cauline leaves). |
Involucres | hemispheric to campanulate, 1.5–2.5 × 1–2.5 cm, thinly arachnoid or glabrate. |
ovoid to broadly cylindric or campanulate, 2–3.5(–4) × 1.5–3 cm, thinly arachnoid. |
Corollas | white to pink, lavender, or purple, 17–25 mm, tubes 5–11 mm, throats 4–7 mm (noticeably wider than tubes), lobes 5–7 mm; style tips 3–4.5 mm. |
pink to purple (white), 25–32 mm, tubes 12–16 mm, throats 7–10 mm, (noticeably wider than tubes), lobes 6–9 mm; style tips 4–6 mm. |
Phyllaries | in 6–10 series, strongly imbricate, green or brownish, ovate or elliptic (outer) to linear-lanceolate (inner), abaxial faces with narrow glutinous ridge; outer and middle appressed, bodies entire, spines abruptly spreading, slender, 1–2(–3) mm; apices of inner often flexuous, flat, attenuate. |
in 10–12 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with narrow glutinous ridge, outer and middle bodies appressed, margins entire, spines abruptly spreading to deflexed, slender, 3–9 mm; spines slender, 3–9 mm; apices of inner phyllaries spreading, narrow, flattened, finely serrulate. |
Heads | few–many, in open corymbiform or paniculiform arrays. |
1–many in corymbiform or paniculiform arrays. |
Cypselae | dark brown, 3–4 mm, apical collars stramineous, 0.5 mm; pappi 17–21 mm (longer bristles shorter than corollas). |
tan to brownish, 4–5 mm, apical collars straw-colored, 0.5–75 mm; pappi 18–25 mm. |
2n | = 24, 26, 28. |
= 20, 21, 22. |
Cirsium nuttallii |
Cirsium discolor |
|
Phenology | Flowering summer (Jun–Aug). | Flowering summer–fall (Jun–Oct). |
Habitat | Roadsides, ditches, woodlands, usually in damp soil | Tallgrass prairies, deciduous woodlands, forest openings, disturbed sites, often in damp soil |
Elevation | 0–100 m (0–300 ft) | 5–800 m (0–2600 ft) |
Distribution |
AL; FL; GA; LA; MS; NC; SC; TX; VA
|
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; NC; NE; NH; NY; OH; PA; RI; SC; SD; TN; VA; VT; WI; WV; MB; ON; QC; SK
|
Discussion | Cirsium nuttallii occurs on the southern coastal plain from southeastern Virginia to southern Florida and west to eastern Louisiana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium discolor is widespread in eastern North America from the prairies of southeastern Saskatchewan, western Minnesota, and Iowa south to northern Louisiana and east across southern Canada to the New England states and the southern Appalachians. It hybridizes with both C. altissimum (discussed thereunder) and C. muticum (G. B. Ownbey 1951b, 1964; W. L. Bloom 1977). Meiosis in first-generation hybrids between C. discolor and C. muticum is usually irregular (Bloom) and most pollen grains are infertile (Ownbey 1951b; Bloom). The presence of a small number of viable cypselae in heads of putative F1 hybrids (Ownbey 1951b) indicates that some F2 hybrids or backcrosses are formed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 119. | FNA vol. 19, p. 112. |
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium |
Sibling taxa | ||
Synonyms | Carduus glaber, Carduus nuttallii | Cnicus discolor, Carduus discolor |
Name authority | de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 651. (1838) | (Muhlenberg ex Willdenow) Sprengel: Syst. Veg. 3: 373. (1826) |
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