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black or Le Conte's thistle, Le Conte's thistle

gray thistle, wavy-leaf thistle, wavyleaf or gray or pasture thistle

Habit Perennials but sometimes appearing biennial, 35–110 cm; taproots, sometimes with root sprouts. Perennials, 20–230 cm; deeply seated runner roots that produce adventitious buds.
Stems

1–few, erect, distal 1/2 nearly naked, loosely arachnoid;

branches 0–5(–10), stiffly ascending.

1–several, erect or ascending, densely gray-tomentose;

branches 0–few, usually above middle, ascending.

Leaves

blades linear to oblong or narrowly elliptic, 15–25 × 1–4 cm, coarsely toothed to shallowly pinnatifid, lobes undivided or coarsely few-toothed, main spines 3–6 mm, abaxial faces often ± glabrate, loosely arachnoid when young, adaxial glabrous or sparingly villous with coarse, multicellular trichomes;

basal sometimes absent at flowering, petiolate;

principal cauline sessile, progressively reduced distally, bases clasping or ± decurrent;

distal cauline few, widely separated, bractlike.

blades elliptic to oblong or ovate, 10–40 × 1–10 cm, margins strongly undulate, coarsely dentate or shallowly to deeply lobed, lobes ascending to spreading, ± triangular, well separated to closely spaced, spinulose and coarsely dentate or usually cleft into 2–3 lanceolate to triangular, often entire-margined, spine-tipped divisions, main spines (yellowish), 2–12+ mm, abaxial densely gray-tomentose, adaxial faces thinly tomentose;

basal sometimes present at flowering, winged-petiolate;

principal cauline becoming sessile and progressively reduced distally, widest at base, bases ± auriculate-clasping to short-decurrent;

distal reduced, spinier.

Peduncles

5–30 cm (elevated above cauline leaves, not subtended by ring of involucre-like bracts).

0–25+ cm.

Involucres

broadly cylindric to campanulate, 2.5–4 × 1.5–4 cm, loosely arachnoid, ± glabrate.

ovoid to hemispheric or broadly campanulate, 2.5–4.5 × 1.5–4.5 cm, loosely arachnoid on phyllary margins or glabrate.

Corollas

pink-purple, 22–45 mm, tubes 10–23 mm, throats 8–14 mm, lobes 7–10 mm;

style tips 4–5 mm.

lavender to pink, purple, or white, 24–50 mm, tubes 12–28 mm, throats 6–14 mm, lobes 6.5–13 mm;

style tips 5–7.5 mm.

Phyllaries

in 6–10 series, imbricate, ovate or lanceolate (outer) to linear-lanceolate (inner), abaxial faces with prominent glutinous ridge, outer and middle tightly appressed, margins spinulose-serrulate, spines ascending, 0.5–2 mm;

apices of inner flat, linear- acuminate.

in 8–12 series, imbricate, ovate to lanceolate (outer) to linear-lanceolate (inner), abaxial faces with prominent glutinous ridge;

outer and middle appressed, spines spreading, 1.5–5 mm;

apices of inner narrow, often flexuous, flat, ± entire, spineless or weakly spiny.

Heads

borne singly or less commonly 2–5(–10) in open, corymbiform arrays.

1–10+, terminal on branches, in leafy, ± corymbiform arrays.

Cypselae

light brown, 5–5.75 mm, apical collars paler than body, ca. 0.75 mm;

pappi 20–40 mm.

light to dark brown, 6–7 mm, bodies and apical collars concolorous, narrow;

pappi 20–38 mm (usually scabridulous).

2n

= 28, 32.

= 26.

Cirsium lecontei

Cirsium undulatum

Phenology Flowering spring–summer (May–Aug). Flowering spring–autumn (May–Oct).
Habitat Sandy pinelands of coastal plain, often in damp soil Mixedgrass prairie, shortgrass prairie, Palouse prairie, sagebrush deserts, pinyon-juniper woodlands, openings in montane coniferous forests, often in disturbed areas
Elevation 0–150 m (0–500 ft) 100–2800 m (300–9200 ft)
Distribution
from FNA
AL; FL; LA; MS; NC; SC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; GA; IA; ID; IL; IN; KS; MI; MN; MO; MT; ND; NE; NM; OK; OR; PA; SD; TX; UT; WA; WI; WY; AB; BC; MB; SK; Mexico (Chihuahua, Coahuila, Durango, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Cirsium lecontei occurs on the southern coastal plain. R. J. Moore and C. Frankton (1969) suggested that it originated as a derivative of ancient hybridization between the ancestors of C. horridulum and C. nuttallii. They further suggested a relationship between C. lecontei and C. grahamii of Arizona and hypothesized an ancient dispersal from the southeastern coastal plain to the western cordillera. Although such relationships are possible, I have seen little support for them in my examination of these taxa. I think it is more likely that C. lecontei, C. horridulum, and C. nuttallii originated from a common stock, and that the resemblances between C. lecontei and C. grahamii are a result of convergence.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cirsium undulatum is widely distributed in the wstern half of North America from the dry plains and plateaus of the Pacific Northwest eastward across the Great Plains to Manitoba and the Dakotas and south to Texas, New Mexico, and northwestern Mexico. It occurs in scattered localities in the Rocky Mountains and northeastern Great Basin region. At least some of the few widely scattered records from the eastern United States are probably introductions. Cirsium undulatum is both widespread and variable. Plants of the Great Plains region tend to be low-growing with a few large heads and elongate corollas. Plants of the Pacific Northwest are usually taller and produce smaller, more numerous heads with shorter corollas. A detailed study of this species might reveal races worthy of recognition as infraspecific taxa.

Wavyleaf thistle is listed by California as a noxious weed. However, most reports of Cirsium undulatum in California are based upon misidentifications of C. canescens. Cirsium undulatum is known to hybridize with C. flodmanii, C. hookerianum, and C. scariosum var. coloradense. J. T. Howell (1960b) reported that C. undulatum was suspected to hybridize with C. brevifolium in the Pacific Northwest.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 114. FNA vol. 19, p. 120.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Synonyms Carduus lecontei Carduus undulatus, C. megacephalum, C. undulatum var. megacephalum
Name authority Torrey & A. Gray: Fl. N. Amer. 2: 458. (1843) (Nuttall) Sprengel: Syst. Veg. 3: 374. (1826)
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