Cirsium lecontei |
Cirsium cymosum |
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black or Le Conte's thistle, Le Conte's thistle |
graygreen thistle, peregrine thistle |
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Habit | Perennials but sometimes appearing biennial, 35–110 cm; taproots, sometimes with root sprouts. | Biennials or perennials, 25–120 cm, pubescence a mixture of fine, non-septate arachnoid trichomes and coarser, septate trichomes, especially along stems and on midveins on abaxial leaf faces, usually ± loose and irregularly deciduous from leaves in age; taprooted. | ||||
Stems | 1–few, erect, distal 1/2 nearly naked, loosely arachnoid; branches 0–5(–10), stiffly ascending. |
usually 1, erect, ± gray-tomentose, sometimes villous with septate trichomes; branches 0–10+, usually arising in distal 1/2, ascending, usually reaching a ± common height. |
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Leaves | blades linear to oblong or narrowly elliptic, 15–25 × 1–4 cm, coarsely toothed to shallowly pinnatifid, lobes undivided or coarsely few-toothed, main spines 3–6 mm, abaxial faces often ± glabrate, loosely arachnoid when young, adaxial glabrous or sparingly villous with coarse, multicellular trichomes; basal sometimes absent at flowering, petiolate; principal cauline sessile, progressively reduced distally, bases clasping or ± decurrent; distal cauline few, widely separated, bractlike. |
blades linear-oblong to oblanceolate or elliptic, 10–30 × 3–7 cm, shallowly to deeply pinnatifid with 3–8 pairs of lobes, longer than 2 cm, lobes well separated, linear to triangular-ovate, dentate to lobed proximally, main spines slender, 2–7 mm, faces green to gray, thinly to densely arachnoid-tomentose with fine, non-septate trichomes, sometimes villous with septate trichomes along veins, usually ± loose and irregularly deciduous from leaves in age; basal often present at flowering, sessile or winged-petiolate; principal cauline mostly in proximal 1/2, winged-petiolate or sessile, bases narrowed, auriculate, veins often prominently raised on abaxial faces; distal sessile, auriculate-clasping or short-decurrent 1–10 mm, progressively reduced becoming bractlike, often unlobed or less deeply divided and sometimes spinier than proximal. |
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Peduncles | 5–30 cm (elevated above cauline leaves, not subtended by ring of involucre-like bracts). |
(0–)2–15 cm. |
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Involucres | broadly cylindric to campanulate, 2.5–4 × 1.5–4 cm, loosely arachnoid, ± glabrate. |
ovoid to hemispheric or campanulate, 2–3 × 1.5–3.5 cm, ± arachnoid-floccose, often glabrate. |
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Corollas | pink-purple, 22–45 mm, tubes 10–23 mm, throats 8–14 mm, lobes 7–10 mm; style tips 4–5 mm. |
creamy white to purplish, 20–31 mm, tubes 8–14 mm, throats 5.5–10 mm, lobes 6–7 mm; style tips 4–6 mm. |
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Phyllaries | in 6–10 series, imbricate, ovate or lanceolate (outer) to linear-lanceolate (inner), abaxial faces with prominent glutinous ridge, outer and middle tightly appressed, margins spinulose-serrulate, spines ascending, 0.5–2 mm; apices of inner flat, linear- acuminate. |
in 8–10 series, subequal to strongly imbricate, green, linear to lanceolate (outer) to linear (inner), entire, abaxial faces with inconspicuous to prominent glutinous ridge; outer and mid bodies loosely spreading to ascending or appressed, apices subappressed to ascending or spreading, flat, spines ascending to spreading, fine, 2–4 mm; apices of inner commonly flexuous or reflexed, narrow, flat, scarious. |
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Heads | borne singly or less commonly 2–5(–10) in open, corymbiform arrays. |
borne singly, terminal on main stem and branches, sometimes also in distal axils, erect, not subtended by well-developed leaves, collectively forming corymbiform or racemiform arrays. |
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Cypselae | light brown, 5–5.75 mm, apical collars paler than body, ca. 0.75 mm; pappi 20–40 mm. |
tan to dark brown, 5–7.5 mm, apical collars not differentiated; pappi 16–25 mm. |
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2n | = 28, 32. |
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Cirsium lecontei |
Cirsium cymosum |
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Phenology | Flowering spring–summer (May–Aug). | |||||
Habitat | Sandy pinelands of coastal plain, often in damp soil | |||||
Elevation | 0–150 m (0–500 ft) | |||||
Distribution |
AL; FL; LA; MS; NC; SC
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CA; ID; MT; NV; OR; WY
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Discussion | Of conservation concern. Cirsium lecontei occurs on the southern coastal plain. R. J. Moore and C. Frankton (1969) suggested that it originated as a derivative of ancient hybridization between the ancestors of C. horridulum and C. nuttallii. They further suggested a relationship between C. lecontei and C. grahamii of Arizona and hypothesized an ancient dispersal from the southeastern coastal plain to the western cordillera. Although such relationships are possible, I have seen little support for them in my examination of these taxa. I think it is more likely that C. lecontei, C. horridulum, and C. nuttallii originated from a common stock, and that the resemblances between C. lecontei and C. grahamii are a result of convergence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Past floras have treated Cirsium cymosum and C. canovirens as separate species. In my examination of these plants across their combined ranges I realized that they are connected by numerous intermediates and that I could find no characters that consistently distinguish them. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 114. | FNA vol. 19, p. 136. | ||||
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Carduus lecontei | Carduus cymosus, C. botrys, C. triacanthum | ||||
Name authority | Torrey & A. Gray: Fl. N. Amer. 2: 458. (1843) | (Greene) J. T. Howell: Amer. Midl. Naturalist 30: 37. (1943) | ||||
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