Cirsium joannae |
Cirsium lecontei |
|
---|---|---|
Joanna's thistle |
black or Le Conte's thistle, Le Conte's thistle |
|
Habit | Perennials, 100–150 cm; caudices taprooted. | Perennials but sometimes appearing biennial, 35–110 cm; taproots, sometimes with root sprouts. |
Stems | 1, fleshy, erect, openly branched in distal 1/2, glabrous; branches on distal stems several, ascending. |
1–few, erect, distal 1/2 nearly naked, loosely arachnoid; branches 0–5(–10), stiffly ascending. |
Leaves | blades oblong, 10–50 × 6–12 cm, margins usually not strongly undulate, coarsely dentate or shallowly to deeply pinnatifid with 10–15 pairs of lobes, teeth or lobes ± closely spaced, not much overlapping, narrowly to broadly triangular, spiny-dentate or larger shallowly 3–5-lobed, main spines 2–12 mm, abaxial faces glabrous, adaxial glabrous; basal present at flowering, petiolate or spiny winged-petiolate; principal cauline many, well distributed, proximally ± winged-petiolate, distally sessile, gradually reduced, less divided, bases auriculate-claping and/or decurrent as spiny wings to 2 cm; distal much reduced. |
blades linear to oblong or narrowly elliptic, 15–25 × 1–4 cm, coarsely toothed to shallowly pinnatifid, lobes undivided or coarsely few-toothed, main spines 3–6 mm, abaxial faces often ± glabrate, loosely arachnoid when young, adaxial glabrous or sparingly villous with coarse, multicellular trichomes; basal sometimes absent at flowering, petiolate; principal cauline sessile, progressively reduced distally, bases clasping or ± decurrent; distal cauline few, widely separated, bractlike. |
Peduncles | 0–2 cm Involucres green, ovoid to campanulate (not including spreading phyllary apices), 2.5–4 × 2.5–3.5 cm, appearing glabrous. |
5–30 cm (elevated above cauline leaves, not subtended by ring of involucre-like bracts). |
Involucres | broadly cylindric to campanulate, 2.5–4 × 1.5–4 cm, loosely arachnoid, ± glabrate. |
|
Corollas | lavender-pink, 20–26 mm, tubes 5–8.5 mm, throats 5.2–8 mm, lobes linear, 9–10 mm; style tips 3.5–4.2 mm, conspicuously exserted beyond corolla lobes. |
pink-purple, 22–45 mm, tubes 10–23 mm, throats 8–14 mm, lobes 7–10 mm; style tips 4–5 mm. |
Phyllaries | in 5–7 series, unequal, outer longer than inner, bases short-appressed, abaxial faces without glutinous ridge, minutely scabridulous, apices green, thick, spreading to curved-ascending, proximally flattened, linear, spines stout, 5–12 mm; outer entire or pinnately spiny, inner entire, scabridulous-ciliolate; apices of inner flexuous, sometimes slightly expanded and minutely erose. |
in 6–10 series, imbricate, ovate or lanceolate (outer) to linear-lanceolate (inner), abaxial faces with prominent glutinous ridge, outer and middle tightly appressed, margins spinulose-serrulate, spines ascending, 0.5–2 mm; apices of inner flat, linear- acuminate. |
Heads | several–many, erect or nodding, usually sessile or short-pedunculate, crowded in subcaptitate or short, spiciform clusters at branch tips, collectively forming open, paniculiform arrays. |
borne singly or less commonly 2–5(–10) in open, corymbiform arrays. |
Cypselae | dark brown, 4.5–5 mm, apical collars not differeniated; pappi ca. 20 mm. |
light brown, 5–5.75 mm, apical collars paler than body, ca. 0.75 mm; pappi 20–40 mm. |
2n | = 28, 32. |
|
Cirsium joannae |
Cirsium lecontei |
|
Phenology | Flowering summer (Aug–Sep). | Flowering spring–summer (May–Aug). |
Habitat | Hanging gardens with Lobelia, Abies, and Adiantum | Sandy pinelands of coastal plain, often in damp soil |
Elevation | 1700 m (5600 ft) | 0–150 m (0–500 ft) |
Distribution |
UT |
AL; FL; LA; MS; NC; SC
|
Discussion | Cirsium joannae is endemic to Zion National Park in southwestern Utah. It apears to be most closely related to C. rydbergii, which occurs in similar habitats in southeastern Utah and northeastern Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Cirsium lecontei occurs on the southern coastal plain. R. J. Moore and C. Frankton (1969) suggested that it originated as a derivative of ancient hybridization between the ancestors of C. horridulum and C. nuttallii. They further suggested a relationship between C. lecontei and C. grahamii of Arizona and hypothesized an ancient dispersal from the southeastern coastal plain to the western cordillera. Although such relationships are possible, I have seen little support for them in my examination of these taxa. I think it is more likely that C. lecontei, C. horridulum, and C. nuttallii originated from a common stock, and that the resemblances between C. lecontei and C. grahamii are a result of convergence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 163. | FNA vol. 19, p. 114. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Carduus lecontei | |
Name authority | S. L. Welsh: in S. L. Welsh et al., Utah Fl. ed. 3, 168. (2003) | Torrey & A. Gray: Fl. N. Amer. 2: 458. (1843) |
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