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Davis' thistle, Greene's thistle, intermountain thistle, Jackson hole thistle

Texas or Texas purple or southern thistle, Texas thistle

Habit Biennials or monocarpic perennials, 20–100 cm; deeply taprooted. Annuals or biennials, 20–200 cm; taprooted.
Stems

1–several, erect, thinly to densely gray-tomentose with fine, non-septate trichomes;

branches 0–many, ascending.

usually single, erect, tomentose to ± glabrate;

branches 0–many, usually restricted to distal part, ascending.

Leaves

blades oblanceolate or elliptic, 10–35 × 1–7 cm, unlobed and spinulose to dentate or deeply pinnatifid, usually 5–8 pairs of lobes, well separated, linear to lance-triangular, spinulose to few toothed or lobed, main spines 2–7 mm, abaxial faces densely gray-tomentose or sometimes ± glabrate, adaxial gray to ± green, thinly tomentose or ± glabrate;

basal sometimes present at flowering, narrowly winged-petiolate;

principal cauline well distributed, gradually reduced, sometimes spinier than basal;

proximal winged-petiolate, mid sessile, bases spiny-winged, decurrent 1–3 cm;

distal becoming bractlike, often unlobed or less deeply divided than proximal.

blades oblong to elliptic, 7–30 × 2–12 cm, unlobed and merely spinulose to irregularly dentate or shallowly to deeply pinnatifid, lobes ± triangular, separated by narrow to wide sinuses, sometimes coarsely dentate or lobed proximally, obtuse to acute, main spines slender to stout, 1–5 mm, abaxial faces arachnoid tomentose, adaxial glabrous or thinly arachnoid;

basal often absent at flowering, petioles slender, ± winged;

cauline progressively reduced, proximal petiolate, mid and distal broadly sessile, bases ± auriculate-clasping or decurrent 1–3 cm;

distalmost linear to lanceolate, bractlike, irregularly dentate or shallowly lobed.

Peduncles

0–25 cm.

slender, 3–30 cm (not overtopped by crowded distal leaves).

Involucres

ovoid or hemispheric to campanulate, 2–3 × 1.5–5 cm, glabrous or loosely floccose to densely arachnoid.

ovoid to hemispheric, 1.5–2 × 1.5–2 cm, thinly arachnoid, glabrate.

Corollas

dull white or faintly lavender-tinged to bright pink-purple, 19–31 mm, tubes 7–13 mm, throats 6.5–9.5 mm, lobes 4–8 mm;

style tips 3.5–7 mm.

white to pink-purple, 20–25 mm, tubes 7–10 mm, throats 6–8 mm (noticeably wider than tubes), lobes 4–7 mm;

style tips 3–4 mm.

Phyllaries

in 6–10 series, strongly imbricate or sometimes subequal, greenish to brown, ovate to linear-lanceolate (outer) to linear (inner), entire, abaxial faces with narrow or scarcely developed glutinous ridge;

outer and mid appressed or apices ascending to spreading, linear, bodies entire, spines ascending to abruptly spreading, usually fine, 2–6 mm;

apices of inner narrow, spine-tipped or spineless.

in 8–10 series, imbricate, green, lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge;

outer and middle appressed, bodies entire, acute, spines spreading, slender, 1–5 mm;

apices of inner often flexuous, flat, scabrid-ciliolate, acuminate.

Heads

1–many, in open corymbiform arrays or crowded near stem tips.

1–many, in openly paniculiform arrays.

Cypselae

brown, 5–8 mm, apical collars not differentiated;

pappi 12–25 mm.

brown, 3–5 mm, apical collars not differentiated;

pappi 15–16 mm.

2n

= 32, 34, 36.

= 22, 23, 24.

Cirsium inamoenum

Cirsium texanum

Phenology Flowering spring–summer (Apr–Jul).
Habitat Roadsides, pastures, fields, shrub-tree savannas
Elevation 0–1000 m (0–3300 ft)
Distribution
from FNA
CA; ID; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; LA; MO; NM; OK; TX; Mexico (Coahuila, Durango, Nuevo León, San Luis Potosí, Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Cirsium inamoenum is a variable complex across the northern Great Basin and adjacent mountains. A. Cronquist (1994) treated this complex as a single species under the name C. subniveum without infraspecific taxa and including taxa that formerly had been assigned to C. utahense (e.g., J. T. Howell 1960b). Some populations consist of small-headed, white-flowered plants with strong involucres and short, appressed phyllaries. Others have larger heads, white or lavender to pink-purple corollas, and phyllaries with longer, ascending to spreading tips. My treatment of this complex as C. inamoenum is similarly inclusive as was Cronquist’s treatment of C. subniveum, except that I believe C. humboldtense, which Cronquist included, is probably a derivative of hybridization between C. subniveum and C. eatonii var. peckii. I have observed such hybrids on the slopes of Steens Mountain in Harney County, Oregon, and the type of C. humboldtense closely resembles some of the introgressants. I have examined several other specimens that are likely the products of hybridization of C. inamoenum with other varieties of C. eatonii.

I have chosen to recognize racial differentiation within Cirsium inamoenum at the rank of variety. The main difference between Cirsium inamoenum var. inamoenum and var. davisii is corolla color. Unfortunately this feature is sometimes difficult to determine on herbarium specimens, and many collectors fail to include corolla color on specimen labels. Some geographic overlap occurs between var. davisii, which has a distribution centered in northeastern Utah, southeastern Idaho, and adjacent southwestern Wyoming, and the more widespread var. inamoenum.

Plants of northeastern Oregon, southeastern Washington, and adjacent western Idaho often have large heads and densely tomentose foliage. These were named Cirsium wallowense by Peck. Similar plants occur sporadically in other portions of the range of Cirsium inamoenum var. inamoenum and I chose not to recognize these northwestern populations as a third variety. Additional study might clarify the relationships of these plants.

Some specimens of Cirsium inamoenum in central Nevada and Utah approach C. neomexicanum. It seems likely that these species have interacted in the past.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cirsium texanum ranges from the Chihuahuan Desert regions of trans-Pecos Texas and adjacent southeastern New Mexico across the plains of Texas and southern Oklahoma to southwestern Arkansas and southwestern Louisiana and south into north-central Mexico. D. S. Correll and M. C. Johnston (1970) suggested hybridization between Cirsium texanum and C. undulatum to explain anomalous specimens in the Edwards Plateau and trans-Pecos regions of western Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas white or pale lavender
var. inamoenum
1. Corollas lavender to rich pink-purple
var. davisii
Source FNA vol. 19, p. 134. FNA vol. 19, p. 119.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. inamoenum var. davisii, C. inamoenum var. inamoenum
Synonyms Carduus inamoenus C. austrinum, C. helleri, C. texanum var. stenolepis
Name authority (Greene) D. J. Keil: Sida 21: 214. (2004) Buckley: Proc. Acad. Nat. Sci. Philadelphia 13: 460. (1862)
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