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Davis' thistle, Greene's thistle, intermountain thistle, Jackson hole thistle

bristly or horrid or yellow or bull thistle, bristly thistle, bull thistle, horrid thistle, yellow thistle

Habit Biennials or monocarpic perennials, 20–100 cm; deeply taprooted. Biennials or perennials, (± fleshy), 15–250 cm; stout taproots and a fascicle of fleshy lateral roots, often perennating by root sprouts.
Stems

1–several, erect, thinly to densely gray-tomentose with fine, non-septate trichomes;

branches 0–many, ascending.

1–several, usually erect, often stout, glabrous to densely tomentose;

branches 0–many, spreading to ascending, short, stout.

Leaves

blades oblanceolate or elliptic, 10–35 × 1–7 cm, unlobed and spinulose to dentate or deeply pinnatifid, usually 5–8 pairs of lobes, well separated, linear to lance-triangular, spinulose to few toothed or lobed, main spines 2–7 mm, abaxial faces densely gray-tomentose or sometimes ± glabrate, adaxial gray to ± green, thinly tomentose or ± glabrate;

basal sometimes present at flowering, narrowly winged-petiolate;

principal cauline well distributed, gradually reduced, sometimes spinier than basal;

proximal winged-petiolate, mid sessile, bases spiny-winged, decurrent 1–3 cm;

distal becoming bractlike, often unlobed or less deeply divided than proximal.

blades linear to oblanceolate or oblong-elliptic, 10–40 × 2–10 cm, unlobed and spiny-dentate to deeply pinnatifid, lobes spiny-dentate or coarsely lobed, main spines stout, 5–30 mm, abaxial faces subglabrous to loosely tomentose, adaxial glabrous to ± densely villous with septate trichomes;

basal present at flowering, spiny winged-petiolate, bases often tapered;

principal cauline sessile, well distributed, often not much reduced distally, bases often ± auriculate-clasping;

distal cauline often spinier than the proximal.

Peduncles

0–25 cm.

0–5 cm.

Involucres

ovoid or hemispheric to campanulate, 2–3 × 1.5–5 cm, glabrous or loosely floccose to densely arachnoid.

hemispheric to campanulate, 3–5 × 3–8 cm.

Corollas

dull white or faintly lavender-tinged to bright pink-purple, 19–31 mm, tubes 7–13 mm, throats 6.5–9.5 mm, lobes 4–8 mm;

style tips 3.5–7 mm.

white to yellow, pink, purple, or red, 30–47 mm, tubes 11–30 mm, throats 6–10 mm, lobes 7–10 mm;

style tips 3–5 mm.

Phyllaries

in 6–10 series, strongly imbricate or sometimes subequal, greenish to brown, ovate to linear-lanceolate (outer) to linear (inner), entire, abaxial faces with narrow or scarcely developed glutinous ridge;

outer and mid appressed or apices ascending to spreading, linear, bodies entire, spines ascending to abruptly spreading, usually fine, 2–6 mm;

apices of inner narrow, spine-tipped or spineless.

in 5–9 series, subequal to imbricate, light green to stramineous, lanceolate to linear, distally often with reddish margins, abaxial faces without glutinous ridge, often ± thinly tomentose, often scabridulous in submarginal bands;

outer and middle appressed-ascending, bodies usually reddish-tinged, margins setulose-ciliolate, apices acuminate, spines 1–2 mm, weak;

apices of inner straight, flat.

Heads

1–many, in open corymbiform arrays or crowded near stem tips.

1–20 in subcapitate to corymbiform arrays (each closely subtended by an involucre-like ring of spiny-margined bracts).

Cypselae

brown, 5–8 mm, apical collars not differentiated;

pappi 12–25 mm.

straw-colored to tan, 4–6 mm, apical collars weakly differentiated;

pappi 25–35 mm.

2n

= 32, 34, 36.

= 32, 33, 34, 35.

Cirsium inamoenum

Cirsium horridulum

Distribution
from FNA
CA; ID; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; FL; GA; LA; MA; MD; ME; MS; NC; NH; NJ; NY; OK; PA; RI; SC; TN; TX; VA; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Cirsium inamoenum is a variable complex across the northern Great Basin and adjacent mountains. A. Cronquist (1994) treated this complex as a single species under the name C. subniveum without infraspecific taxa and including taxa that formerly had been assigned to C. utahense (e.g., J. T. Howell 1960b). Some populations consist of small-headed, white-flowered plants with strong involucres and short, appressed phyllaries. Others have larger heads, white or lavender to pink-purple corollas, and phyllaries with longer, ascending to spreading tips. My treatment of this complex as C. inamoenum is similarly inclusive as was Cronquist’s treatment of C. subniveum, except that I believe C. humboldtense, which Cronquist included, is probably a derivative of hybridization between C. subniveum and C. eatonii var. peckii. I have observed such hybrids on the slopes of Steens Mountain in Harney County, Oregon, and the type of C. humboldtense closely resembles some of the introgressants. I have examined several other specimens that are likely the products of hybridization of C. inamoenum with other varieties of C. eatonii.

I have chosen to recognize racial differentiation within Cirsium inamoenum at the rank of variety. The main difference between Cirsium inamoenum var. inamoenum and var. davisii is corolla color. Unfortunately this feature is sometimes difficult to determine on herbarium specimens, and many collectors fail to include corolla color on specimen labels. Some geographic overlap occurs between var. davisii, which has a distribution centered in northeastern Utah, southeastern Idaho, and adjacent southwestern Wyoming, and the more widespread var. inamoenum.

Plants of northeastern Oregon, southeastern Washington, and adjacent western Idaho often have large heads and densely tomentose foliage. These were named Cirsium wallowense by Peck. Similar plants occur sporadically in other portions of the range of Cirsium inamoenum var. inamoenum and I chose not to recognize these northwestern populations as a third variety. Additional study might clarify the relationships of these plants.

Some specimens of Cirsium inamoenum in central Nevada and Utah approach C. neomexicanum. It seems likely that these species have interacted in the past.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (3 in the flora).

Although several variants have been given taxonomic recognition as species, these seem at most races. Flower color varies greatly, sometimes within populations and sometimes on a populational or regional basis. Herbarium specimens are sometimes difficult to assign to variety.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas white or pale lavender
var. inamoenum
1. Corollas lavender to rich pink-purple
var. davisii
1. Stems densely tomentose; involucres ± densely tomentose
var. horridulum
1. Stems glabrous or sparsely tomentose; involucres glabrous
→ 2
2. Leaves shallowly to deeply pinnatifid, main spines 10–30 mm
var. megacanthum
2. Leaves spinose-dentate to shallowly pinnatifid, main spines mostly 5–10 mm
var. vittatum
Source FNA vol. 19, p. 134. FNA vol. 19, p. 114.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. inamoenum var. davisii, C. inamoenum var. inamoenum
C. horridulum var. horridulum, C. horridulum var. megacanthum, C. horridulum var. vittatum
Synonyms Carduus inamoenus
Name authority (Greene) D. J. Keil: Sida 21: 214. (2004) Michaux: Fl. Bor.-Amer. 2: 90. (1803)
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