Flora of North America species comparison

usually 1 and erect, less commonly several and ascending, simple to sparingly short-branched in distal 1/2, variably villous with jointed trichomes, and/or finely arachnoid, or ± glabrate;

branches on distal stems 0–many, short, ascending.

1–few, erect, closely gray-tomentose;

branches 0–few, ascending.

blades linear-oblong to elliptic, 5–25 × 1–8 cm, subentire to coarsely dentate or deeply pinnatifid, lobes lance-oblong to broadly triangular, spinulose to spiny-dentate or shallowly lobed, main spines 2–10 mm, abaxial faces usually ± densely gray- or white-tomentose with felted arachnoid trichomes, ± villous to tomentose along major veins with septate trichomes, sometimes glabrous or glabrate, adaxial ± green, glabrous to thinly arachnoid, often ± villous or tomentose with septate trichomes;

basal often present at flowering, spiny winged-petiolate or sessile;

principal cauline well distributed, proximally winged-petiolate, distally sessile, gradually reduced, bases sometimes short-decurrent;

distal ± reduced, often narrower than proximal, sometimes with non-pigmented bases, sometimes pectinately spiny.

blades lanceolate to narrowly elliptic, 10–25 × 1–4 cm, unlobed and merely spinulose or pinnately lobed about halfway to midveins, often terminal lobes long-tapered, lobes short, lanceolate to triangular, entire to few toothed or lobed, well separated by wide, U-shaped sinuses, main spines slender, 2–5 mm, abaxial faces gray-tomentose, adaxial green, glabrous to thinly tomentose;

basal usually present at flowering, winged-petiolate;

principal cauline winged-petiolate proximally, mid and distal sessile, progressively reduced distally, bases not or scarcely decurrent, sometimes distal weakly clasping;

distalmost often reduced to lanceolate or linear, long-acuminate bracts.

0–8+ cm.

0–10 cm.

(green or often purplish), broadly ovoid, 2–3.3 × 1.5–4 cm, loosely to densely villous with septate trichomes to tomentose and/or arachnoid.

hemispheric to subcylindric, 1.5–2.2 × 1.5–2.5 cm, thinly floccose-tomentose or glabrate.

white, ochroleucous, or occasionally pink, 20–28 mm, tubes 10–13 mm, throats 6.5–9 mm, lobes 5–7 mm;

style tips 3–5.5 mm.

white or pink to pale purple, 20–28 mm, tubes 9–14 mm, throats 5–7.5 mm, lobes 5–10 mm;

style tips 2.5–6 mm.

in 4–8 series, imbricate to subequal, bases short-appressed, entire, abaxial faces with or without narrow glutinous ridge, apices stiffly spreading to ascending, linear, long, plane, spines straight, slender, 3–5 mm;

apices of inner flexuous, sometimes expanded and erose.

in 6–9 series, imbricate, pale green with darker apices, brownish when dry, lanceolate (outer) to linear-lanceolate (inner), margins of outer entire, abaxial faces with narrow glutinous ridge;

outer and middle appressed proximally, apices spreading to ascending, bodies entire or rarely spinulose, spines slender, 3–7 mm;

apices of inner often dark purple or blackish, flexuous, scarious, entire to pectinate-fringed, tapered or expanded.

1–many, borne singly or crowded in spiciform, racemiform, subcapitate, or sometimes more openly branched corymbiform arrays.

1–6, borne singly or few at branch tips in corymbiform arrays.

dark brown, 5–6.5 mm, apical collars not differentiated;

pappi 18–22 mm.

stramineous with brownish streaks, 6.5–7 mm, apical collars colored like body, narrow;

pappi 15–20 mm.

= 34.

= 28.