Flora of North America species comparison

usually 1 and erect, less commonly several and ascending, simple to sparingly short-branched in distal 1/2, variably villous with jointed trichomes, and/or finely arachnoid, or ± glabrate;

branches on distal stems 0–many, short, ascending.

usually single, erect, tomentose to ± glabrate;

branches 0–many, usually restricted to distal part, ascending.

blades linear-oblong to elliptic, 5–25 × 1–8 cm, subentire to coarsely dentate or deeply pinnatifid, lobes lance-oblong to broadly triangular, spinulose to spiny-dentate or shallowly lobed, main spines 2–10 mm, abaxial faces usually ± densely gray- or white-tomentose with felted arachnoid trichomes, ± villous to tomentose along major veins with septate trichomes, sometimes glabrous or glabrate, adaxial ± green, glabrous to thinly arachnoid, often ± villous or tomentose with septate trichomes;

basal often present at flowering, spiny winged-petiolate or sessile;

principal cauline well distributed, proximally winged-petiolate, distally sessile, gradually reduced, bases sometimes short-decurrent;

distal ± reduced, often narrower than proximal, sometimes with non-pigmented bases, sometimes pectinately spiny.

blades oblong to elliptic, 7–30 × 2–12 cm, unlobed and merely spinulose to irregularly dentate or shallowly to deeply pinnatifid, lobes ± triangular, separated by narrow to wide sinuses, sometimes coarsely dentate or lobed proximally, obtuse to acute, main spines slender to stout, 1–5 mm, abaxial faces arachnoid tomentose, adaxial glabrous or thinly arachnoid;

basal often absent at flowering, petioles slender, ± winged;

cauline progressively reduced, proximal petiolate, mid and distal broadly sessile, bases ± auriculate-clasping or decurrent 1–3 cm;

distalmost linear to lanceolate, bractlike, irregularly dentate or shallowly lobed.

0–8+ cm.

slender, 3–30 cm (not overtopped by crowded distal leaves).

(green or often purplish), broadly ovoid, 2–3.3 × 1.5–4 cm, loosely to densely villous with septate trichomes to tomentose and/or arachnoid.

ovoid to hemispheric, 1.5–2 × 1.5–2 cm, thinly arachnoid, glabrate.

white, ochroleucous, or occasionally pink, 20–28 mm, tubes 10–13 mm, throats 6.5–9 mm, lobes 5–7 mm;

style tips 3–5.5 mm.

white to pink-purple, 20–25 mm, tubes 7–10 mm, throats 6–8 mm (noticeably wider than tubes), lobes 4–7 mm;

style tips 3–4 mm.

in 4–8 series, imbricate to subequal, bases short-appressed, entire, abaxial faces with or without narrow glutinous ridge, apices stiffly spreading to ascending, linear, long, plane, spines straight, slender, 3–5 mm;

apices of inner flexuous, sometimes expanded and erose.

in 8–10 series, imbricate, green, lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge;

outer and middle appressed, bodies entire, acute, spines spreading, slender, 1–5 mm;

apices of inner often flexuous, flat, scabrid-ciliolate, acuminate.

1–many, borne singly or crowded in spiciform, racemiform, subcapitate, or sometimes more openly branched corymbiform arrays.

1–many, in openly paniculiform arrays.

dark brown, 5–6.5 mm, apical collars not differentiated;

pappi 18–22 mm.

brown, 3–5 mm, apical collars not differentiated;

pappi 15–16 mm.

= 34.

= 22, 23, 24.