Flora of North America species comparison

usually 1 and erect, less commonly several and ascending, simple to sparingly short-branched in distal 1/2, variably villous with jointed trichomes, and/or finely arachnoid, or ± glabrate;

branches on distal stems 0–many, short, ascending.

1 or few, erect, densely gray-tomentose;

branches 0 to several, ascending to spreading.

blades linear-oblong to elliptic, 5–25 × 1–8 cm, subentire to coarsely dentate or deeply pinnatifid, lobes lance-oblong to broadly triangular, spinulose to spiny-dentate or shallowly lobed, main spines 2–10 mm, abaxial faces usually ± densely gray- or white-tomentose with felted arachnoid trichomes, ± villous to tomentose along major veins with septate trichomes, sometimes glabrous or glabrate, adaxial ± green, glabrous to thinly arachnoid, often ± villous or tomentose with septate trichomes;

basal often present at flowering, spiny winged-petiolate or sessile;

principal cauline well distributed, proximally winged-petiolate, distally sessile, gradually reduced, bases sometimes short-decurrent;

distal ± reduced, often narrower than proximal, sometimes with non-pigmented bases, sometimes pectinately spiny.

blades elliptic to obovate, 10–30 × 8–14 cm, deeply divided nearly to midveins, lobes ascending to spreading, linear, remote, margins revolute, entire or minutely spinulose, main spines 1–2 mm, faces gray-tomentose, more densely so abaxially;

basal present or withered at flowering, petiolate;

principal cauline well distributed, bases decurrent as linear-lobed to spiny wings 1–3 cm;

distal cauline well developed.

0–8+ cm.

0–5 cm.

(green or often purplish), broadly ovoid, 2–3.3 × 1.5–4 cm, loosely to densely villous with septate trichomes to tomentose and/or arachnoid.

ovoid to campanulate, 2–3 × 2–3 cm, loosely arachnoid on phyllary margins or glabrate.

white, ochroleucous, or occasionally pink, 20–28 mm, tubes 10–13 mm, throats 6.5–9 mm, lobes 5–7 mm;

style tips 3–5.5 mm.

dull white or pinkish-tinged (rarely rich purple), 20–30 mm, tubes 8.5–15 mm, throats 4.5–10 mm, lobes 3–8 mm;

style tips 3.5–5.5 mm.

in 4–8 series, imbricate to subequal, bases short-appressed, entire, abaxial faces with or without narrow glutinous ridge, apices stiffly spreading to ascending, linear, long, plane, spines straight, slender, 3–5 mm;

apices of inner flexuous, sometimes expanded and erose.

in 6–8 series, imbricate, ovate-lanceolate (outer) to linear-lanceolate (inner), abaxial faces with narrow glutinous ridge;

outer and middle appressed, acute, spines ascending to spreading, slender, 1–2(–3) mm;

apices of inner often flexuous, flattened, spineless, scabrid.

1–many, borne singly or crowded in spiciform, racemiform, subcapitate, or sometimes more openly branched corymbiform arrays.

1–20+ in corymbiform arrays.

dark brown, 5–6.5 mm, apical collars not differentiated;

pappi 18–22 mm.

light brown, sometimes with darker streaks, 6–7.5 mm, apical collars lighter colored, very narrow;

pappi 15–30 mm, usually noticeably shorter than corolla.

= 34.

= 34.