Flora of North America species comparison

usually 1 and erect, less commonly several and ascending, simple to sparingly short-branched in distal 1/2, variably villous with jointed trichomes, and/or finely arachnoid, or ± glabrate;

branches on distal stems 0–many, short, ascending.

1–several, erect, gray- or white-tomentose;

branches 0–few, ascending.

blades linear-oblong to elliptic, 5–25 × 1–8 cm, subentire to coarsely dentate or deeply pinnatifid, lobes lance-oblong to broadly triangular, spinulose to spiny-dentate or shallowly lobed, main spines 2–10 mm, abaxial faces usually ± densely gray- or white-tomentose with felted arachnoid trichomes, ± villous to tomentose along major veins with septate trichomes, sometimes glabrous or glabrate, adaxial ± green, glabrous to thinly arachnoid, often ± villous or tomentose with septate trichomes;

basal often present at flowering, spiny winged-petiolate or sessile;

principal cauline well distributed, proximally winged-petiolate, distally sessile, gradually reduced, bases sometimes short-decurrent;

distal ± reduced, often narrower than proximal, sometimes with non-pigmented bases, sometimes pectinately spiny.

blades oblong-oblanceolate to narrowly elliptic, 4–40 × 1–10 cm, bases usually not decurrent, finely spiny-toothed and undivided to coarsely toothed or deeply pinnatifid, lobes broadly triangular to linear-lanceolate, often revolute-margined, main spines 1–7 mm, abaxial faces white-tomentose, adaxial faces green, thinly tomentose, ± glabrate;

basal usually absent or withered at flowering, winged petiolate;

principal cauline proximally winged-petiolate, distally sessile, well distributed, gradually reduced, bases usually not decurrent;

distal cauline well developed.

0–8+ cm.

0–5 cm (elevated above distal leaves).

(green or often purplish), broadly ovoid, 2–3.3 × 1.5–4 cm, loosely to densely villous with septate trichomes to tomentose and/or arachnoid.

ovoid to broadly campanulate, 2–3.5 × 2.5–3.5 cm, thinly arachnoid.

white, ochroleucous, or occasionally pink, 20–28 mm, tubes 10–13 mm, throats 6.5–9 mm, lobes 5–7 mm;

style tips 3–5.5 mm.

purple (white), 23–36 mm, tubes 12–15 mm, throats 6–8.5 mm, lobes 5–9 mm;

style tips 4–7 mm.

in 4–8 series, imbricate to subequal, bases short-appressed, entire, abaxial faces with or without narrow glutinous ridge, apices stiffly spreading to ascending, linear, long, plane, spines straight, slender, 3–5 mm;

apices of inner flexuous, sometimes expanded and erose.

in 7–12 series, strongly imbricate, greenish with subapical darker central zone, ovate or lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge;

outer and middle entire, bodies appressed, entire, acute, spines abruptly spreading, slender, 2–4 mm;

apices of inner spreading, flexuous, narrow, flattened, finely serrulate, ± scabrous.

1–many, borne singly or crowded in spiciform, racemiform, subcapitate, or sometimes more openly branched corymbiform arrays.

erect, borne singly and terminal on main stem and branches, or few in corymbiform arrays from distal axils (not subtended by ring of spiny-margined bracts).

dark brown, 5–6.5 mm, apical collars not differentiated;

pappi 18–22 mm.

light brown, 3–5 mm, apical collars stramineous, 0.5–1 mm;

pappi (white or tawny) 20–30 mm.

= 34.

= 22, 24.