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Graham's thistle

alcove thistle, Rydberg's thistle

Habit Biennials, 50–100 cm; taproots slender and fascicles of thick fibrous roots. Perennials, 100–300 cm; caudices and taproots, spreading by creeping roots.
Stems

1, erect, thinly arachnoid and/or puberulent to short-pilose, sometimes ± glabrate;

branches 0–4, ascending.

1–several, erect or ascending to lax and hanging, glabrous or thinly tomentose;

branches 0–few, ascending.

Leaves

blades oblanceolate to oblong-elliptic, 20–30 × 3–8 cm, spinulose and otherwise entire or coarsely dentate to deeply pinnatifid, lobes entire or coarsely few toothed or lobed, main spines slender, 3–6 mm, abaxial ± persistently gray-tomentose, sometimes pilose along veins, adaxial faces thinly arachnoid and ± glabrate;

basal often present at flowering, sessile or narrowly winged-petiolate;

principal cauline gradually winged-petiolate or sessile, reduced distally, bases sometimes clasping or short-decurrent;

distal cauline ascending, becoming bractlike, narrow, lobed or not.

blades elliptic, 30–90+ × 10–40 cm, 1–2 times pinnately lobed, lobes linear to ovate, strongly undulate, main spines slender, 5–15 mm, faces often glaucous, glabrous or thinly tomentose and soon glabrescent;

basal present at flowering, petiolate or winged-petiolate;

proximal cauline winged-petiolate;

mid sessile, much reduced, less deeply lobed, bases clasping, short-decurrent 0–2 cm;

distal linear or lanceolate, bractlike, very spiny.

Peduncles

10–30 cm.

0.5–6 cm.

Involucres

hemispheric, 2–3 × 2–4 cm, thinly arachnoid or glabrous.

hemispheric, 1.4–2 × 1–2 cm, phyllary margins thinly tomentose or glabrate.

Corollas

deep purple, 22–30 mm, tubes 13–18 mm, throats 4–5 mm, lobes 5–8 mm;

style tips 4–4.5 mm.

dull white to pink or purple, 16–20 mm, tubes 7–8.5 mm, throats 4–6.5 mm, lobes 4.5–6 mm;

style tips 2.5 mm.

Phyllaries

in ca. 8 series, imbricate, proximally brownish, distally dark purplish, lanceolate to linear, margins of outer hispidulous-ciliolate, spiny fringed, pinnately spiny or with scarious appendages, abaxial faces with prominent, glutinous ridge;

outer and middle appressed or only apices spreading, bodies minutely spinulose-denticulate, spines erect to ascending, 1.5–2.5 mm;

apices of inner phyllaries often flexuous, flat, scabridulous.

in 5–8 series, strongly imbricate, (green, drying green or light brown), ovate to lance-oblong, abaxial faces with or without poorly developed glutinous ridge;

outer and mid bases appressed, margins entire, not scabridulous-ciliolate, apices spreading or reflexed, green to brownish, lance-ovate, elongate, flattened, spines slender, 3–25 mm;

apices of inner straight, entire.

Heads

1–5.

few–many, erect or nodding in clusters at tips of distal branches in paniculiform arrays, not closely subtended by clustered leafy bracts.

Cypselae

tan with dark speckles to dark purplish brown, 4–5.5 mm, apical collars not differently colored;

pappi 13–18 mm.

gray or brown, 3.7–4.5 mm, apical collars not differentiated;

pappi 10–15 mm.

2n

= 32 (Mexico).

= 34.

Cirsium grahamii

Cirsium rydbergii

Phenology Flowering Jul–Sep. Flowering spring–summer (May–Sep).
Habitat Oak woodlands, coniferous forests, meadows, often in damp soil Hanging gardens, seeps, stream banks
Elevation 1400–2600 m (4600–8500 ft) 1000–1500 m (3300–4900 ft)
Distribution
from FNA
AZ; NM; Mexico (Chihuahua, Durango, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; UT
[BONAP county map]
Discussion

Cirsium grahamii occurs in the mountains of southeastern Arizona and southwestern New Mexico. It forms hybrid swarms with C. parryi and C. scariosum var. coloradense in the White Mountains of Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cirsium rydbergii is endemic to the Colorado Plateau of northern Arizona and southeastern Utah.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 124. FNA vol. 19, p. 162.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Synonyms C. lactucinum
Name authority A. Gray: Smithsonian Contr. Knowl. 5(6): 102. (1853) Petrak: Beih. Bot. Centralbl. 35(2): 315. (1917)
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