Cirsium grahamii |
Cirsium engelmannii |
|
---|---|---|
Graham's thistle |
blackland thistle, Engelmann's thistle |
|
Habit | Biennials, 50–100 cm; taproots slender and fascicles of thick fibrous roots. | Biennials or monocarpic perennials, 40–200 cm; taproots and clusters of coarse fibrous roots that often have tuberlike thickenings. |
Stems | 1, erect, thinly arachnoid and/or puberulent to short-pilose, sometimes ± glabrate; branches 0–4, ascending. |
single, erect, often branched above middle, thinly arachnoid-tomentose, ± glabrate; branches few, ascending. |
Leaves | blades oblanceolate to oblong-elliptic, 20–30 × 3–8 cm, spinulose and otherwise entire or coarsely dentate to deeply pinnatifid, lobes entire or coarsely few toothed or lobed, main spines slender, 3–6 mm, abaxial ± persistently gray-tomentose, sometimes pilose along veins, adaxial faces thinly arachnoid and ± glabrate; basal often present at flowering, sessile or narrowly winged-petiolate; principal cauline gradually winged-petiolate or sessile, reduced distally, bases sometimes clasping or short-decurrent; distal cauline ascending, becoming bractlike, narrow, lobed or not. |
blades elliptic or ovate, 5–20 × 1–10 cm, usually deeply pinnatifid, lobes narrowly to broadly triangular, sinuses broad, rounded (basal and distal cauline sometimes less divided, lobes linear-lanceolate), margins revolute, spreading, entire or spinulose to remotely few-toothed or sharply lobed, main spines slender, 1–5 mm, abaxial faces white-tomentose, adaxial green, villous with septate trichomes or glabrate; basal usually absent at flowering, winged-petiolate, bases tapered; principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping; distal reduced, widely separated, distalmost bractlike. |
Peduncles | 10–30 cm. |
2–20+ cm, essentially naked with much reduced bracts. |
Involucres | hemispheric, 2–3 × 2–4 cm, thinly arachnoid or glabrous. |
ovoid to broadly cylindric or campanulate, 2.5–3.5 × 2–3 cm, thinly arachnoid. |
Corollas | deep purple, 22–30 mm, tubes 13–18 mm, throats 4–5 mm, lobes 5–8 mm; style tips 4–4.5 mm. |
pink to purple (white), 32–38 mm, tubes 15–20 mm, throats 6–9 mm, lobes 8–11 mm; style tips 5–6 mm. |
Phyllaries | in ca. 8 series, imbricate, proximally brownish, distally dark purplish, lanceolate to linear, margins of outer hispidulous-ciliolate, spiny fringed, pinnately spiny or with scarious appendages, abaxial faces with prominent, glutinous ridge; outer and middle appressed or only apices spreading, bodies minutely spinulose-denticulate, spines erect to ascending, 1.5–2.5 mm; apices of inner phyllaries often flexuous, flat, scabridulous. |
in 10–12 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with narrow glutinous ridge; outer and middle entire, bodies appressed, spines abruptly spreading to deflexed, slender, 2–4 mm; apices of inner phyllaries narrow, flexuous, flattened, entire or finely erose. |
Heads | 1–5. |
1–10+, borne at tips of main stem and branches. |
Cypselae | tan with dark speckles to dark purplish brown, 4–5.5 mm, apical collars not differently colored; pappi 13–18 mm. |
brown, 5–6 mm, apical collars yellow, ca. 1 mm; pappi 25–30 mm. |
2n | = 32 (Mexico). |
= 18 (as C. terrae-nigrae), 20 + 1B. |
Cirsium grahamii |
Cirsium engelmannii |
|
Phenology | Flowering Jul–Sep. | Flowering spring–summer (May–Jul). |
Habitat | Oak woodlands, coniferous forests, meadows, often in damp soil | Tallgrass prairies, old fields, roadsides, oak savannas, forest edges, in calcareous clay or rarely sandy soils |
Elevation | 1400–2600 m (4600–8500 ft) | 50–200 m (200–700 ft) |
Distribution |
AZ; NM; Mexico (Chihuahua, Durango, Sonora)
|
LA; OK; TX |
Discussion | Cirsium grahamii occurs in the mountains of southeastern Arizona and southwestern New Mexico. It forms hybrid swarms with C. parryi and C. scariosum var. coloradense in the White Mountains of Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium engelmannii occurs mostly in the blackland prairies of eastern Texas. It ranges northward into southeastern Oklahoma and eastward to northwestern Arkansas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 124. | FNA vol. 19, p. 117. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. virginianum var. filipendulum, C. terrae-nigrae | |
Name authority | A. Gray: Smithsonian Contr. Knowl. 5(6): 102. (1853) | Rydberg: Fl. Rocky Mts., 1069. (1917) |
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