Cirsium flodmanii |
Cirsium joannae |
|
---|---|---|
chardon de Flodman, Flodman's thistle, prairie thistle |
Joanna's thistle |
|
Habit | Perennials 30–140 cm; horizontal runner roots that produce root sprouts. | Perennials, 100–150 cm; caudices taprooted. |
Stems | 1–several, erect, gray- or white-tomentose; branches 0–few, ascending. |
1, fleshy, erect, openly branched in distal 1/2, glabrous; branches on distal stems several, ascending. |
Leaves | blades oblong-oblanceolate to narrowly elliptic, 4–40 × 1–10 cm, bases usually not decurrent, finely spiny-toothed and undivided to coarsely toothed or deeply pinnatifid, lobes broadly triangular to linear-lanceolate, often revolute-margined, main spines 1–7 mm, abaxial faces white-tomentose, adaxial faces green, thinly tomentose, ± glabrate; basal usually absent or withered at flowering, winged petiolate; principal cauline proximally winged-petiolate, distally sessile, well distributed, gradually reduced, bases usually not decurrent; distal cauline well developed. |
blades oblong, 10–50 × 6–12 cm, margins usually not strongly undulate, coarsely dentate or shallowly to deeply pinnatifid with 10–15 pairs of lobes, teeth or lobes ± closely spaced, not much overlapping, narrowly to broadly triangular, spiny-dentate or larger shallowly 3–5-lobed, main spines 2–12 mm, abaxial faces glabrous, adaxial glabrous; basal present at flowering, petiolate or spiny winged-petiolate; principal cauline many, well distributed, proximally ± winged-petiolate, distally sessile, gradually reduced, less divided, bases auriculate-claping and/or decurrent as spiny wings to 2 cm; distal much reduced. |
Peduncles | 0–5 cm (elevated above distal leaves). |
0–2 cm Involucres green, ovoid to campanulate (not including spreading phyllary apices), 2.5–4 × 2.5–3.5 cm, appearing glabrous. |
Involucres | ovoid to broadly campanulate, 2–3.5 × 2.5–3.5 cm, thinly arachnoid. |
|
Corollas | purple (white), 23–36 mm, tubes 12–15 mm, throats 6–8.5 mm, lobes 5–9 mm; style tips 4–7 mm. |
lavender-pink, 20–26 mm, tubes 5–8.5 mm, throats 5.2–8 mm, lobes linear, 9–10 mm; style tips 3.5–4.2 mm, conspicuously exserted beyond corolla lobes. |
Phyllaries | in 7–12 series, strongly imbricate, greenish with subapical darker central zone, ovate or lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge; outer and middle entire, bodies appressed, entire, acute, spines abruptly spreading, slender, 2–4 mm; apices of inner spreading, flexuous, narrow, flattened, finely serrulate, ± scabrous. |
in 5–7 series, unequal, outer longer than inner, bases short-appressed, abaxial faces without glutinous ridge, minutely scabridulous, apices green, thick, spreading to curved-ascending, proximally flattened, linear, spines stout, 5–12 mm; outer entire or pinnately spiny, inner entire, scabridulous-ciliolate; apices of inner flexuous, sometimes slightly expanded and minutely erose. |
Heads | erect, borne singly and terminal on main stem and branches, or few in corymbiform arrays from distal axils (not subtended by ring of spiny-margined bracts). |
several–many, erect or nodding, usually sessile or short-pedunculate, crowded in subcaptitate or short, spiciform clusters at branch tips, collectively forming open, paniculiform arrays. |
Cypselae | light brown, 3–5 mm, apical collars stramineous, 0.5–1 mm; pappi (white or tawny) 20–30 mm. |
dark brown, 4.5–5 mm, apical collars not differeniated; pappi ca. 20 mm. |
2n | = 22, 24. |
|
Cirsium flodmanii |
Cirsium joannae |
|
Phenology | Flowering summer (Jun–Sep). | Flowering summer (Aug–Sep). |
Habitat | Tallgrass, mixedgrass, shortgrass prairies, meadows, pastures, often in damp soil | Hanging gardens with Lobelia, Abies, and Adiantum |
Elevation | 100–2400 m (300–7900 ft) | 1700 m (5600 ft) |
Distribution |
CO; IA; IL; KS; MI; MN; MT; ND; NE; SD; WI; WY; AB; MB; ON; QC; SK
|
UT |
Discussion | Cirsium flodmanii ranges from Saskatchewan and Alberta south through the northern Great Plains and intermountain valleys of Montana and Wyoming to northeastern Colorado and east through the prairies to Minnesota and Iowa, and in widely scattered locations eastward to northern Illinois, southern Wisconsin, southern Ontario, and southern Quebec. It is known to hybridize with C. muticum and C. undulatum. Hybrids between C. flodmanii and C. undulatum are highly sterile with numerous meiotic irregularities (S. Dabydeen 1987). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium joannae is endemic to Zion National Park in southwestern Utah. It apears to be most closely related to C. rydbergii, which occurs in similar habitats in southeastern Utah and northeastern Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 120. | FNA vol. 19, p. 163. |
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium |
Sibling taxa | ||
Synonyms | Carduus flodmanii, C. oblanceolatum | |
Name authority | (Rydberg) Arthur: Torreya 12: 34. (1912) | S. L. Welsh: in S. L. Welsh et al., Utah Fl. ed. 3, 168. (2003) |
Web links |