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chardon de Flodman, Flodman's thistle, prairie thistle

Graham's thistle

Habit Perennials 30–140 cm; horizontal runner roots that produce root sprouts. Biennials, 50–100 cm; taproots slender and fascicles of thick fibrous roots.
Stems

1–several, erect, gray- or white-tomentose;

branches 0–few, ascending.

1, erect, thinly arachnoid and/or puberulent to short-pilose, sometimes ± glabrate;

branches 0–4, ascending.

Leaves

blades oblong-oblanceolate to narrowly elliptic, 4–40 × 1–10 cm, bases usually not decurrent, finely spiny-toothed and undivided to coarsely toothed or deeply pinnatifid, lobes broadly triangular to linear-lanceolate, often revolute-margined, main spines 1–7 mm, abaxial faces white-tomentose, adaxial faces green, thinly tomentose, ± glabrate;

basal usually absent or withered at flowering, winged petiolate;

principal cauline proximally winged-petiolate, distally sessile, well distributed, gradually reduced, bases usually not decurrent;

distal cauline well developed.

blades oblanceolate to oblong-elliptic, 20–30 × 3–8 cm, spinulose and otherwise entire or coarsely dentate to deeply pinnatifid, lobes entire or coarsely few toothed or lobed, main spines slender, 3–6 mm, abaxial ± persistently gray-tomentose, sometimes pilose along veins, adaxial faces thinly arachnoid and ± glabrate;

basal often present at flowering, sessile or narrowly winged-petiolate;

principal cauline gradually winged-petiolate or sessile, reduced distally, bases sometimes clasping or short-decurrent;

distal cauline ascending, becoming bractlike, narrow, lobed or not.

Peduncles

0–5 cm (elevated above distal leaves).

10–30 cm.

Involucres

ovoid to broadly campanulate, 2–3.5 × 2.5–3.5 cm, thinly arachnoid.

hemispheric, 2–3 × 2–4 cm, thinly arachnoid or glabrous.

Corollas

purple (white), 23–36 mm, tubes 12–15 mm, throats 6–8.5 mm, lobes 5–9 mm;

style tips 4–7 mm.

deep purple, 22–30 mm, tubes 13–18 mm, throats 4–5 mm, lobes 5–8 mm;

style tips 4–4.5 mm.

Phyllaries

in 7–12 series, strongly imbricate, greenish with subapical darker central zone, ovate or lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge;

outer and middle entire, bodies appressed, entire, acute, spines abruptly spreading, slender, 2–4 mm;

apices of inner spreading, flexuous, narrow, flattened, finely serrulate, ± scabrous.

in ca. 8 series, imbricate, proximally brownish, distally dark purplish, lanceolate to linear, margins of outer hispidulous-ciliolate, spiny fringed, pinnately spiny or with scarious appendages, abaxial faces with prominent, glutinous ridge;

outer and middle appressed or only apices spreading, bodies minutely spinulose-denticulate, spines erect to ascending, 1.5–2.5 mm;

apices of inner phyllaries often flexuous, flat, scabridulous.

Heads

erect, borne singly and terminal on main stem and branches, or few in corymbiform arrays from distal axils (not subtended by ring of spiny-margined bracts).

1–5.

Cypselae

light brown, 3–5 mm, apical collars stramineous, 0.5–1 mm;

pappi (white or tawny) 20–30 mm.

tan with dark speckles to dark purplish brown, 4–5.5 mm, apical collars not differently colored;

pappi 13–18 mm.

2n

= 22, 24.

= 32 (Mexico).

Cirsium flodmanii

Cirsium grahamii

Phenology Flowering summer (Jun–Sep). Flowering Jul–Sep.
Habitat Tallgrass, mixedgrass, shortgrass prairies, meadows, pastures, often in damp soil Oak woodlands, coniferous forests, meadows, often in damp soil
Elevation 100–2400 m (300–7900 ft) 1400–2600 m (4600–8500 ft)
Distribution
from FNA
CO; IA; IL; KS; MI; MN; MT; ND; NE; SD; WI; WY; AB; MB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; Mexico (Chihuahua, Durango, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Cirsium flodmanii ranges from Saskatchewan and Alberta south through the northern Great Plains and intermountain valleys of Montana and Wyoming to northeastern Colorado and east through the prairies to Minnesota and Iowa, and in widely scattered locations eastward to northern Illinois, southern Wisconsin, southern Ontario, and southern Quebec. It is known to hybridize with C. muticum and C. undulatum. Hybrids between C. flodmanii and C. undulatum are highly sterile with numerous meiotic irregularities (S. Dabydeen 1987).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cirsium grahamii occurs in the mountains of southeastern Arizona and southwestern New Mexico. It forms hybrid swarms with C. parryi and C. scariosum var. coloradense in the White Mountains of Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 120. FNA vol. 19, p. 124.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Synonyms Carduus flodmanii, C. oblanceolatum
Name authority (Rydberg) Arthur: Torreya 12: 34. (1912) A. Gray: Smithsonian Contr. Knowl. 5(6): 102. (1853)
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