Flora of North America species comparison

single, erect, often branched above middle, thinly arachnoid-tomentose, ± glabrate;

branches few, ascending.

1–20+, erect or ascending, densely gray-tomentose with non-septate trichomes;

branches 0 or few, usually in distal 1/2, ascending.

blades elliptic or ovate, 5–20 × 1–10 cm, usually deeply pinnatifid, lobes narrowly to broadly triangular, sinuses broad, rounded (basal and distal cauline sometimes less divided, lobes linear-lanceolate), margins revolute, spreading, entire or spinulose to remotely few-toothed or sharply lobed, main spines slender, 1–5 mm, abaxial faces white-tomentose, adaxial green, villous with septate trichomes or glabrate;

basal usually absent at flowering, winged-petiolate, bases tapered;

principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping;

distal reduced, widely separated, distalmost bractlike.

blades oblong to narrowly elliptic, 10–30 × 2–8 cm, strongly undulate, margins coarsely dentate or shallowly to deeply pinnatifid with 8–15 pairs of lobes 0.5–2 cm, often revolute, lobes ± triangular, closely spaced, spreading, spinose-dentate and cleft into 2–5 spine-tipped divisions, main spines 5–20 mm, yellowish, abaxial faces densely white-tomentose, adaxial thinly gray-tomentose;

basal usually present at flowering, winged-petiolate;

principal cauline sessile, progressively reduced distally, bases ± auriculate to long-decurrent as spiny wings;

distal cauline usually much reduced, less lobed.

2–20+ cm, essentially naked with much reduced bracts.

0–4 cm.

ovoid to broadly cylindric or campanulate, 2.5–3.5 × 2–3 cm, thinly arachnoid.

ovoid to hemispheric or broadly campanulate, 2.5–4.5 × 2.5–4.5 cm in first-formed heads, often smaller in later ones, loosely arachnoid on phyllary margins or glabrate.

pink to purple (white), 32–38 mm, tubes 15–20 mm, throats 6–9 mm, lobes 8–11 mm;

style tips 5–6 mm.

white or pale lavender to purple, pink, or red, 25–45 mm, tubes 8–25 mm, throats 6–17 mm, lobes 6–15 mm;

style tips 2–8 mm.

in 10–12 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with narrow glutinous ridge;

outer and middle entire, bodies appressed, spines abruptly spreading to deflexed, slender, 2–4 mm;

apices of inner phyllaries narrow, flexuous, flattened, entire or finely erose.

in 5–10 series, imbricate, ovate (outer) to linear-lanceolate (inner), margins entire, abaxial faces with narrow glutinous ridge;

outer and middle appressed, spines spreading, 3–12 mm;

apices of inner often flexuous, expanded and flat, scabrid-margined, sometimes erose, spineless.

1–10+, borne at tips of main stem and branches.

1–few, in leafy, ± corymbiform arrays.

brown, 5–6 mm, apical collars yellow, ca. 1 mm;

pappi 25–30 mm.

light brown, sometimes with lighter or darker streaks, 6–9 mm, apical collars colored like the body, narrow;

pappi (white or tawny), 20–40 mm, usually noticeably shorter than corolla.

= 18 (as C. terrae-nigrae), 20 + 1B.

= 30, 31, 32, 34.