Cirsium engelmannii |
Cirsium hydrophilum |
|||||
---|---|---|---|---|---|---|
blackland thistle, Engelmann's thistle |
Mt. Tamalpais thistle, Suisun thistle |
|||||
Habit | Biennials or monocarpic perennials, 40–200 cm; taproots and clusters of coarse fibrous roots that often have tuberlike thickenings. | Biennials or monocarpic perennials, 100–220 cm; taprooted. | ||||
Stems | single, erect, often branched above middle, thinly arachnoid-tomentose, ± glabrate; branches few, ascending. |
1–several, erect, (hollow), openly branched distally or throughout, thinly arachnoid with fine, non-septate trichomes, glabrate. |
||||
Leaves | blades elliptic or ovate, 5–20 × 1–10 cm, usually deeply pinnatifid, lobes narrowly to broadly triangular, sinuses broad, rounded (basal and distal cauline sometimes less divided, lobes linear-lanceolate), margins revolute, spreading, entire or spinulose to remotely few-toothed or sharply lobed, main spines slender, 1–5 mm, abaxial faces white-tomentose, adaxial green, villous with septate trichomes or glabrate; basal usually absent at flowering, winged-petiolate, bases tapered; principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping; distal reduced, widely separated, distalmost bractlike. |
blades elliptic to broadly oblanceolate, 10–40+ cm, pinnatifid 1/2–2/3 distance to midveins, larger usually with broad sinuses, lobes broad, few lobed or dentate, main spines 2–9 mm, abaxial faces ± gray-tomentose, sometimes ± glabrate, adaxial thinly arachnoid-tomentose, soon glabrescent; basal present or withered at flowering, winged-petiolate; principal cauline sessile, progressively reduced distally, bases auriculate-clasping or shortly decurrent; distal cauline reduced, bractlike, often spinier than proximal. |
||||
Peduncles | 2–20+ cm, essentially naked with much reduced bracts. |
0–10+ cm. |
||||
Involucres | ovoid to broadly cylindric or campanulate, 2.5–3.5 × 2–3 cm, thinly arachnoid. |
ovoid to campanulate, 1.5–2.5 × 1.5–3 cm, thinly arachnoid, glabrate. |
||||
Corollas | pink to purple (white), 32–38 mm, tubes 15–20 mm, throats 6–9 mm, lobes 8–11 mm; style tips 5–6 mm. |
pale rose-purple, 18–23 mm, tubes 8–10 mm, throats 5–6 mm, lobes 5–7 mm; style tips 3.5–4.5 mm. |
||||
Phyllaries | in 10–12 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with narrow glutinous ridge; outer and middle entire, bodies appressed, spines abruptly spreading to deflexed, slender, 2–4 mm; apices of inner phyllaries narrow, flexuous, flattened, entire or finely erose. |
in 6–9 series, imbricate, dark green to brownish, lanceolate (outer) to linear (inner), abaxial faces with narrow glutinous ridge; outer and middle appressed, apices spreading, finely serrulate, spines slender, 1–2 mm; apices of inner erect, ± flexuous. |
||||
Heads | 1–10+, borne at tips of main stem and branches. |
borne singly or few at branch tips, sometimes subtended by clustered, ± leafy bracts, collectively forming ± open, many-headed paniculiform arrays. |
||||
Cypselae | brown, 5–6 mm, apical collars yellow, ca. 1 mm; pappi 25–30 mm. |
dark brown to black, 4–5 mm, collars very narrow, stramineous; pappi ca. 15 mm. |
||||
2n | = 18 (as C. terrae-nigrae), 20 + 1B. |
= 32. |
||||
Cirsium engelmannii |
Cirsium hydrophilum |
|||||
Phenology | Flowering spring–summer (May–Jul). | |||||
Habitat | Tallgrass prairies, old fields, roadsides, oak savannas, forest edges, in calcareous clay or rarely sandy soils | |||||
Elevation | 50–200 m (200–700 ft) | |||||
Distribution |
LA; OK; TX |
CA
|
||||
Discussion | Cirsium engelmannii occurs mostly in the blackland prairies of eastern Texas. It ranges northward into southeastern Oklahoma and eastward to northwestern Arkansas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||
Key |
|
|||||
Source | FNA vol. 19, p. 117. | FNA vol. 19, p. 132. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | C. virginianum var. filipendulum, C. terrae-nigrae | Carduus hydrophilus, C. vaseyi var. hydrophilum | ||||
Name authority | Rydberg: Fl. Rocky Mts., 1069. (1917) | (Greene) Jepson: Fl. W. Calif., 507. (1901) | ||||
Web links |