Cirsium engelmannii |
Cirsium brevifolium |
|
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blackland thistle, Engelmann's thistle |
Palouse thistle |
|
Habit | Biennials or monocarpic perennials, 40–200 cm; taproots and clusters of coarse fibrous roots that often have tuberlike thickenings. | Perennials, 25–120 cm; taproots with horizontal root sprouts. |
Stems | single, erect, often branched above middle, thinly arachnoid-tomentose, ± glabrate; branches few, ascending. |
1–several, erect, thinly gray-tomentose with fine, non-septate trichomes; branches 0–many, ascending. |
Leaves | blades elliptic or ovate, 5–20 × 1–10 cm, usually deeply pinnatifid, lobes narrowly to broadly triangular, sinuses broad, rounded (basal and distal cauline sometimes less divided, lobes linear-lanceolate), margins revolute, spreading, entire or spinulose to remotely few-toothed or sharply lobed, main spines slender, 1–5 mm, abaxial faces white-tomentose, adaxial green, villous with septate trichomes or glabrate; basal usually absent at flowering, winged-petiolate, bases tapered; principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping; distal reduced, widely separated, distalmost bractlike. |
oblanceolate or elliptic, 15–45 × 2–10 cm, unlobed and merely spinulose to dentate or deeply pinnatifid, lobes well separated, linear to triangular-ovate, merely spinulose to few toothed or lobed near base, margins often revolute, main spines 2–3(–6) mm, abaxial faces densely gray-tomentose, adaxial green, thinly tomentose or ± glabrate; basal often present at flowering, narrowly winged-petiolate; principal cauline well distributed, gradually reduced distally, bases of proximal cauline winged-petiolate or sessile, bases of distal cauline expanded and ± clasping, margins sometimes spinier than those of proximal; distalmost cauline becoming bractlike, often unlobed or less deeply divided than proximal. |
Peduncles | 2–20+ cm, essentially naked with much reduced bracts. |
1–8 cm. |
Involucres | ovoid to broadly cylindric or campanulate, 2.5–3.5 × 2–3 cm, thinly arachnoid. |
hemispheric to campanulate, 2.5–3.5 × 2–4 cm, glabrous or loosely floccose. |
Corollas | pink to purple (white), 32–38 mm, tubes 15–20 mm, throats 6–9 mm, lobes 8–11 mm; style tips 5–6 mm. |
creamy white, rarely lavender-tinged, 22–28 mm, tubes 8–13 mm, throats 7–11 mm, lobes 4–6 mm; style tips 5–6 mm. |
Phyllaries | in 10–12 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with narrow glutinous ridge; outer and middle entire, bodies appressed, spines abruptly spreading to deflexed, slender, 2–4 mm; apices of inner phyllaries narrow, flexuous, flattened, entire or finely erose. |
in 6–10 series, strongly imbricate, greenish to brown, ovate to lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge; outer and middle appressed, bodies entire, spines abruptly spreading, fine, 2–3(–5) mm; apices of inner commonly flexuous or reflexed, flat, scarious. |
Heads | 1–10+, borne at tips of main stem and branches. |
borne singly and terminal on main stems and branches or few from distal axils in corymbiform or paniculiform arrays. |
Cypselae | brown, 5–6 mm, apical collars yellow, ca. 1 mm; pappi 25–30 mm. |
brown, 5–6 mm, apical collars yellowish, 0.5–1 mm; pappi 18–22 mm. |
2n | = 18 (as C. terrae-nigrae), 20 + 1B. |
= 22, 26. |
Cirsium engelmannii |
Cirsium brevifolium |
|
Phenology | Flowering spring–summer (May–Jul). | Flowering summer (Jun–Oct). |
Habitat | Tallgrass prairies, old fields, roadsides, oak savannas, forest edges, in calcareous clay or rarely sandy soils | Palouse prairie |
Elevation | 50–200 m (200–700 ft) | 600–1300 m (2000–4300 ft) |
Distribution |
LA; OK; TX |
ID; OR; WA
|
Discussion | Cirsium engelmannii occurs mostly in the blackland prairies of eastern Texas. It ranges northward into southeastern Oklahoma and eastward to northwestern Arkansas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium brevifolium occurs in the Palouse prairie region of eastern Washington, eastern Oregon, and western Idaho. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 117. | FNA vol. 19, p. 124. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. virginianum var. filipendulum, C. terrae-nigrae | C. palousense |
Name authority | Rydberg: Fl. Rocky Mts., 1069. (1917) | Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 421. (1841) |
Web links |