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blackland thistle, Engelmann's thistle

Habit Biennials or monocarpic perennials, 40–200 cm; taproots and clusters of coarse fibrous roots that often have tuberlike thickenings.
Stems

single, erect, often branched above middle, thinly arachnoid-tomentose, ± glabrate;

branches few, ascending.

Leaves

blades elliptic or ovate, 5–20 × 1–10 cm, usually deeply pinnatifid, lobes narrowly to broadly triangular, sinuses broad, rounded (basal and distal cauline sometimes less divided, lobes linear-lanceolate), margins revolute, spreading, entire or spinulose to remotely few-toothed or sharply lobed, main spines slender, 1–5 mm, abaxial faces white-tomentose, adaxial green, villous with septate trichomes or glabrate;

basal usually absent at flowering, winged-petiolate, bases tapered;

principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping;

distal reduced, widely separated, distalmost bractlike.

Peduncles

2–20+ cm, essentially naked with much reduced bracts.

Involucres

ovoid to broadly cylindric or campanulate, 2.5–3.5 × 2–3 cm, thinly arachnoid.

Corollas

pink to purple (white), 32–38 mm, tubes 15–20 mm, throats 6–9 mm, lobes 8–11 mm;

style tips 5–6 mm.

Phyllaries

in 10–12 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with narrow glutinous ridge;

outer and middle entire, bodies appressed, spines abruptly spreading to deflexed, slender, 2–4 mm;

apices of inner phyllaries narrow, flexuous, flattened, entire or finely erose.

Heads

1–10+, borne at tips of main stem and branches.

Cypselae

brown, 5–6 mm, apical collars yellow, ca. 1 mm;

pappi 25–30 mm.

2n

= 18 (as C. terrae-nigrae), 20 + 1B.

Cirsium engelmannii

Phenology Flowering spring–summer (May–Jul).
Habitat Tallgrass prairies, old fields, roadsides, oak savannas, forest edges, in calcareous clay or rarely sandy soils
Elevation 50–200 m (200–700 ft)
Distribution
from FNA
LA; OK; TX
[BONAP county map]
Discussion

Cirsium engelmannii occurs mostly in the blackland prairies of eastern Texas. It ranges northward into southeastern Oklahoma and eastward to northwestern Arkansas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 117.
Parent taxa Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Synonyms C. virginianum var. filipendulum, C. terrae-nigrae
Name authority Rydberg: Fl. Rocky Mts., 1069. (1917)
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