Cirsium crassicaule |
Cirsium muticum |
|
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slough thistle |
chardon mutique, dunce-nettle, horsetops, swamp thistle |
|
Habit | Annuals or biennials, (60–)100–300 cm; taprooted. | Biennials, 30–230 cm; taproots fleshy. |
Stems | usually 1, erect, stout, (hollow, 2–10 cm diam. at base), openly branched distally, thinly arachnoid, villous with jointed trichomes, at least proximally. |
single, erect, villous with septate trichomes or glabrate, distally sometimes thinly tomentose; branches few–many, ascending. |
Leaves | blades elliptic to broadly oblanceolate, 15–70 × 30–150+ cm, flat, pinnatifid 1/2–2/3 distance to midvein, larger usually with broad sinuses, lobes broad, few lobed or dentate, main spines 3–8 mm, abaxial faces gray-tomentose, adaxial thinly arachnoid-tomentose, sometimes midveins with jointed trichomes; basal present or withered at flowering, winged-petiolate; principal cauline sessile, progressively reduced distally, bases clasping or short-decurrent 1–2 cm; distal cauline reduced, becoming bractlike, sometimes spinier than proximal. |
ovate to broadly elliptic or obovate, 15–55 × 4–20 cm, deeply pinnatifid, to 7/8 to midribs, lobes linear to lanceolate, acute to acuminate, irregularly few toothed or lobed, main spines 2–3 mm, abaxial faces thinly tomentose or glabrate, villous with septate trichomes on the veins, adaxial faces thinly pilose; basal usually absent at flowering, petioles spiny-winged, bases tapered; principal cauline petiolate or sessile, gradually reduced distally, bases sometimes ± clasping, not decurrent; distal cauline bractlike with narrowly linear lobes, often spinier than the proximal. |
Peduncles | 0–15 cm. |
0–15 cm (sometimes overtopped by distal cauline leaves, not subtended by involucre-like ring of bracts). |
Involucres | ovoid to campanulate, 1.5–3 × 1.5–3 cm, ± glabrous. |
ovoid to broadly cylindric or campanulate, 1.7–3 × 1–3 cm, arachnoid. |
Corollas | pale rose-purple (white), 19–26 mm, tubes 9–12 mm, throats 4–6 mm, lobes 5–9 mm; style tips 3.5–4.5 mm. |
lavender or purple (white), 16–32 mm, tubes 7–15 mm, throats 4.5–10 mm (noticeably wider than tubes), lobes 4–8 mm; style tips 3.5–5 mm. |
Phyllaries | in 5–7 series, weakly unequal, dark green to brownish, lanceolate (outer) to linear (inner), abaxial faces without (or with very obscure) glutinous ridge; outer and middle appressed or apices spreading, at least outer irregularly spiny-fringed, finely serrulate, spines slender, 3–5 mm; apices of inner erect, abaxial faces gray-tomentose, ± twisted. |
in 8–12 series, strongly imbricate, dull green with darker subapical patch, ovate (outer) to linear-lanceolate (inner), abaxial faces with narrow glutinous ridge, outer and middle appressed, bodies minutely spinulose, apices obtuse to acute, spines erect (sometimes appearing as spreading in dry specimens), 0–0.5 mm; apices of inner phyllaries straight or ± flexuous, flattened. |
Heads | 1–several at branch tips, closely subtended by clustered leafy bracts or not, collectively forming open, corymbiform or paniculiform arrays. |
1–many in ± open corymbiform or paniculiform arrays. |
Cypselae | dark brown, 5–5.5 mm, collars narrow, ± stramineous; pappi 15–20 mm. |
dark brown, 4.5–5.5 mm, apical collars yellow, 0.3 mm; pappi 12–20 mm. |
2n | = 32. |
= 20, 21, 22, 23, 30. |
Cirsium crassicaule |
Cirsium muticum |
|
Phenology | Flowering spring (Apr–Jun). | Flowering summer (Jul–Sep). |
Habitat | Freshwater marshes, canal banks | Wet soil in meadows, prairies, marshes, swamps, bogs, open woods |
Elevation | 5–100 m (0–300 ft) | 0–1500+ m (0–4900+ ft) |
Distribution |
CA |
AL; AR; CT; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; NC; ND; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; MB; NB; NL; NS; ON; PE; QC; SK; SPM
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Discussion | Cirsium crassicaule is known only from a few sites in the San Joaquin Valley. Some populations are threatened by habitat modification and development pressures. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium muticum is very widely distributed across the eastern half of North America from the prairies of southeastern Saskatchewan across southern Canada to Newfoundland and south in the United States from North Dakota and Maine to southeastern Texas and northern Florida. It is more common in the northern half of this range and extends from the coastal plain to the Appalachian highlands. The widely scattered populations in coastal lowlands in the southern United States may be relicts of the glacial distribution of the species. Cirsium muticum is known to hybridize with C. discolor (discussed thereunder) and C. flodmanii. Draining and modification of wetlands have affected populations of C. muticum in some areas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 132. | FNA vol. 19, p. 113. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Carduus crassicaulis | Carduus muticus, C. muticum var. monticola |
Name authority | (Greene) Jepson: Fl. W. Calif., 506. (1901) | Michaux: Fl. Bor.-Amer. 2: 89. (1803) |
Web links |