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Fish Lake thistle, fringe thistle

slough thistle

Habit Biennials or monocarpic or polycarpic perennials, 20–100 cm; taproots sometimes with branched caudices. Annuals or biennials, (60–)100–300 cm; taprooted.
Stems

1–several, erect or ascending, glabrous or thinly arachnoid-tomentose;

branches 0–10+, slender, usually arising in distal 1/2, ascending.

usually 1, erect, stout, (hollow, 2–10 cm diam. at base), openly branched distally, thinly arachnoid, villous with jointed trichomes, at least proximally.

Leaves

blades oblong to oblanceolate or elliptic, 5–40 × 3–11 cm, unlobed and merely spinulose-dentate or more commonly regularly deeply pinnatifid, lobes well separated to crowded, linear to triangular-ovate, ascending-spreading to retrorse, merely spinulose to coarsely dentate or proximally few-lobed, main spines 2–5(–7) mm, slender, abaxial faces green to gray, glabrous or thinly to densely arachnoid-tomentose, sometimes glabrate, often villous with septate trichomes along veins, adaxial green, glabrous;

basal usually present at flowering, sessile or petiolate;

principal cauline well distributed, proximal usually winged-petiolate, mid sessile, decurrent as spiny wings 1–3 cm;

distal cauline ± reduced.

blades elliptic to broadly oblanceolate, 15–70 × 30–150+ cm, flat, pinnatifid 1/2–2/3 distance to midvein, larger usually with broad sinuses, lobes broad, few lobed or dentate, main spines 3–8 mm, abaxial faces gray-tomentose, adaxial thinly arachnoid-tomentose, sometimes midveins with jointed trichomes;

basal present or withered at flowering, winged-petiolate;

principal cauline sessile, progressively reduced distally, bases clasping or short-decurrent 1–2 cm;

distal cauline reduced, becoming bractlike, sometimes spinier than proximal.

Peduncles

0–30 cm.

0–15 cm.

Involucres

ovoid to campanulate, 1.5–3 × 1–3 cm, glabrous to thinly arachnoid-tomentose and/or villous-ciliate, with long septate trichomes connecting adjacent phyllaries.

ovoid to campanulate, 1.5–3 × 1.5–3 cm, ± glabrous.

Corollas

creamy white to pale pinkish, 16–20 mm, tubes 6.5–9 mm, throats 4–7.5 mm, lobes 4–6 mm;

style tips 3.5–5 mm.

pale rose-purple (white), 19–26 mm, tubes 9–12 mm, throats 4–6 mm, lobes 5–9 mm;

style tips 3.5–4.5 mm.

Phyllaries

in 5–6 series, imbricate or subequal, outer green or with maroon to dark brown subapical patch or appendage, linear to ovate, abaxial faces with narrow glutinous ridge that may be concealed by trichomes;

outer and middle with bases appressed, apical appendages erect or ascending, ovate to linear-lanceolate or acicular, entire or spinulose to broadly expanded, scarious, and erose-dentate, apical appendages, spines erect or ascending, 1–5 mm, ± flattened;

apices of inner sometimes flexuous or reflexed, narrow, flat, entire or ± expanded, scarious and lacerate-dentate.

in 5–7 series, weakly unequal, dark green to brownish, lanceolate (outer) to linear (inner), abaxial faces without (or with very obscure) glutinous ridge;

outer and middle appressed or apices spreading, at least outer irregularly spiny-fringed, finely serrulate, spines slender, 3–5 mm;

apices of inner erect, abaxial faces gray-tomentose, ± twisted.

Heads

few–many, borne singly or clustered in corymbiform, paniculiform, or racemiform arrays at tips of main stem and branches, sometimes also in distal axils not closely subtended by clustered leafy bracts.

1–several at branch tips, closely subtended by clustered leafy bracts or not, collectively forming open, corymbiform or paniculiform arrays.

Cypselae

tan to dark brown, 5–6 mm, apical collars not or scarcely differentiated;

pappi 14–16 mm.

dark brown, 5–5.5 mm, collars narrow, ± stramineous;

pappi 15–20 mm.

2n

= 32.

Cirsium clavatum

Cirsium crassicaule

Phenology Flowering spring (Apr–Jun).
Habitat Freshwater marshes, canal banks
Elevation 5–100 m (0–300 ft)
Distribution
from FNA
CO; UT; WY; Central Rocky Mountains
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Cirsium clavatum is a polymorphic and variable species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cirsium crassicaule is known only from a few sites in the San Joaquin Valley. Some populations are threatened by habitat modification and development pressures.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres densely villous or tomentose with long, septate trichomes connecting adjacent phyllaries
var. osterhoutii
1. Involucres glabrous or thinly arachnoid-tomentose with fine, non-septate trichomes
→ 2
2. Some or all of the phyllaries usually with dilated, scarious, erose to fringed appendages; mostly Colorado and Wyoming
var. americanum
2. Phyllaries usually entire; w Colorado and Utah
var. clavatum
Source FNA vol. 19, p. 126. FNA vol. 19, p. 132.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. clavatum var. americanum, C. clavatum var. clavatum, C. clavatum var. osterhoutii
Synonyms Cnicus clavatus Carduus crassicaulis
Name authority (M. E. Jones) Petrak: Beih. Bot. Centralbl. 35(2): 310. (1917) (Greene) Jepson: Fl. W. Calif., 506. (1901)
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