Cirsium clavatum |
Cirsium crassicaule |
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Fish Lake thistle, fringe thistle |
slough thistle |
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Habit | Biennials or monocarpic or polycarpic perennials, 20–100 cm; taproots sometimes with branched caudices. | Annuals or biennials, (60–)100–300 cm; taprooted. | ||||||||
Stems | 1–several, erect or ascending, glabrous or thinly arachnoid-tomentose; branches 0–10+, slender, usually arising in distal 1/2, ascending. |
usually 1, erect, stout, (hollow, 2–10 cm diam. at base), openly branched distally, thinly arachnoid, villous with jointed trichomes, at least proximally. |
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Leaves | blades oblong to oblanceolate or elliptic, 5–40 × 3–11 cm, unlobed and merely spinulose-dentate or more commonly regularly deeply pinnatifid, lobes well separated to crowded, linear to triangular-ovate, ascending-spreading to retrorse, merely spinulose to coarsely dentate or proximally few-lobed, main spines 2–5(–7) mm, slender, abaxial faces green to gray, glabrous or thinly to densely arachnoid-tomentose, sometimes glabrate, often villous with septate trichomes along veins, adaxial green, glabrous; basal usually present at flowering, sessile or petiolate; principal cauline well distributed, proximal usually winged-petiolate, mid sessile, decurrent as spiny wings 1–3 cm; distal cauline ± reduced. |
blades elliptic to broadly oblanceolate, 15–70 × 30–150+ cm, flat, pinnatifid 1/2–2/3 distance to midvein, larger usually with broad sinuses, lobes broad, few lobed or dentate, main spines 3–8 mm, abaxial faces gray-tomentose, adaxial thinly arachnoid-tomentose, sometimes midveins with jointed trichomes; basal present or withered at flowering, winged-petiolate; principal cauline sessile, progressively reduced distally, bases clasping or short-decurrent 1–2 cm; distal cauline reduced, becoming bractlike, sometimes spinier than proximal. |
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Peduncles | 0–30 cm. |
0–15 cm. |
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Involucres | ovoid to campanulate, 1.5–3 × 1–3 cm, glabrous to thinly arachnoid-tomentose and/or villous-ciliate, with long septate trichomes connecting adjacent phyllaries. |
ovoid to campanulate, 1.5–3 × 1.5–3 cm, ± glabrous. |
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Corollas | creamy white to pale pinkish, 16–20 mm, tubes 6.5–9 mm, throats 4–7.5 mm, lobes 4–6 mm; style tips 3.5–5 mm. |
pale rose-purple (white), 19–26 mm, tubes 9–12 mm, throats 4–6 mm, lobes 5–9 mm; style tips 3.5–4.5 mm. |
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Phyllaries | in 5–6 series, imbricate or subequal, outer green or with maroon to dark brown subapical patch or appendage, linear to ovate, abaxial faces with narrow glutinous ridge that may be concealed by trichomes; outer and middle with bases appressed, apical appendages erect or ascending, ovate to linear-lanceolate or acicular, entire or spinulose to broadly expanded, scarious, and erose-dentate, apical appendages, spines erect or ascending, 1–5 mm, ± flattened; apices of inner sometimes flexuous or reflexed, narrow, flat, entire or ± expanded, scarious and lacerate-dentate. |
in 5–7 series, weakly unequal, dark green to brownish, lanceolate (outer) to linear (inner), abaxial faces without (or with very obscure) glutinous ridge; outer and middle appressed or apices spreading, at least outer irregularly spiny-fringed, finely serrulate, spines slender, 3–5 mm; apices of inner erect, abaxial faces gray-tomentose, ± twisted. |
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Heads | few–many, borne singly or clustered in corymbiform, paniculiform, or racemiform arrays at tips of main stem and branches, sometimes also in distal axils not closely subtended by clustered leafy bracts. |
1–several at branch tips, closely subtended by clustered leafy bracts or not, collectively forming open, corymbiform or paniculiform arrays. |
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Cypselae | tan to dark brown, 5–6 mm, apical collars not or scarcely differentiated; pappi 14–16 mm. |
dark brown, 5–5.5 mm, collars narrow, ± stramineous; pappi 15–20 mm. |
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2n | = 32. |
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Cirsium clavatum |
Cirsium crassicaule |
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Phenology | Flowering spring (Apr–Jun). | |||||||||
Habitat | Freshwater marshes, canal banks | |||||||||
Elevation | 5–100 m (0–300 ft) | |||||||||
Distribution |
CO; UT; WY; Central Rocky Mountains
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CA |
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Discussion | Varieties 3 (3 in the flora). Cirsium clavatum is a polymorphic and variable species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium crassicaule is known only from a few sites in the San Joaquin Valley. Some populations are threatened by habitat modification and development pressures. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 126. | FNA vol. 19, p. 132. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Cnicus clavatus | Carduus crassicaulis | ||||||||
Name authority | (M. E. Jones) Petrak: Beih. Bot. Centralbl. 35(2): 310. (1917) | (Greene) Jepson: Fl. W. Calif., 506. (1901) | ||||||||
Web links |