Cirsium canescens |
Cirsium pitcheri |
|
---|---|---|
Platte thistle, prairie thistle |
dune thistle, Pitcher's thistle, sand-dune thistle |
|
Habit | Biennials or monocarpic perennials, 20–100 cm; taproots long. | Biennials or short-lived monocarpic perennials, 20–100 cm; taproots long. |
Stems | usually 1, erect, ± densely gray-tomentose with fine, non-septate trichomes; branches 0 or few, usually above middle in distal 1/2, ascending. |
1 or few, erect, densely gray-tomentose; branches 0 to several, ascending to spreading. |
Leaves | blades oblong to elliptic or obovate, 10–25(–40) × 2–6(–12) cm, coarsely dentate or shallowly lobed to deeply pinnatifid, lobes well separated, triangular to linear or oblong, often revolute-margined, ascending to spreading, spinulose to spinose-dentate, main spines 2–3(–10) mm, faces gray-tomentose, more densely abaxially, sometimes glabrate adaxially; basal usually present at flowering, winged-petiolate; principal cauline progressively reduced distally, bases decurrent as spiny wings 1–5 cm, sometimes with expanded auricles; distal cauline usually much reduced, less lobed. |
blades elliptic to obovate, 10–30 × 8–14 cm, deeply divided nearly to midveins, lobes ascending to spreading, linear, remote, margins revolute, entire or minutely spinulose, main spines 1–2 mm, faces gray-tomentose, more densely so abaxially; basal present or withered at flowering, petiolate; principal cauline well distributed, bases decurrent as linear-lobed to spiny wings 1–3 cm; distal cauline well developed. |
Peduncles | 0–10 cm. |
0–5 cm. |
Involucres | hemispheric to broadly campanulate, usually truncate or indented at base, 3–4 × 2.5–4 cm in first-formed heads, often smaller (1.5–2 cm) in later ones, loosely arachnoid on phyllary margins or glabrate. |
ovoid to campanulate, 2–3 × 2–3 cm, loosely arachnoid on phyllary margins or glabrate. |
Corollas | dull white or lavender-tinged, 20–35 mm, tubes 10–17 mm, throats 6–11 mm, lobes 4–9 mm; style tips 5–8 mm. |
dull white or pinkish-tinged (rarely rich purple), 20–30 mm, tubes 8.5–15 mm, throats 4.5–10 mm, lobes 3–8 mm; style tips 3.5–5.5 mm. |
Phyllaries | in 6–9 series, imbricate, ovate-lanceolate (outer) to linear-lanceolate (inner), abaxial faces with prominent glutinous ridge; bodies of outer and middle appressed, acute, spines ascending to spreading, 2–4(–8) mm; apices of inner expanded and flat, often twisted, scabrid-margined, and erose, spineless. |
in 6–8 series, imbricate, ovate-lanceolate (outer) to linear-lanceolate (inner), abaxial faces with narrow glutinous ridge; outer and middle appressed, acute, spines ascending to spreading, slender, 1–2(–3) mm; apices of inner often flexuous, flattened, spineless, scabrid. |
Heads | 1–10+, terminal on branches or in distal axils, in openly corymbiform to racemiform arrays. |
1–20+ in corymbiform arrays. |
Cypselae | light brown, 5–7 mm, sometimes with darker streaks, apical collar very narrow, lighter colored; pappi 18–30 mm, usually noticeably shorter than corolla. |
light brown, sometimes with darker streaks, 6–7.5 mm, apical collars lighter colored, very narrow; pappi 15–30 mm, usually noticeably shorter than corolla. |
2n | = 34, 36. |
= 34. |
Cirsium canescens |
Cirsium pitcheri |
|
Phenology | Flowering spring–summer (May–Aug). | Flowering spring–summer (May–Sep). |
Habitat | Sandy or gravelly soils in short-grass prairie, often in disturbed areas, mountain meadows, grassy slopes in montane coniferous forests | Sand dunes and beaches |
Elevation | 1100–3800 m (3600–12500 ft) | 180–200 m (600–700 ft) |
Distribution |
CA; CO; MO; MT; NE; NV; SD; WY
|
IL; IN; MI; WI; ON
|
Discussion | Cirsium canescens grows in the northern Great Plains from eastern Montana and Wyoming to eastern Colorado and Nebraska; an upland race occurs in the Rocky Mountains of eastern Colorado. It has been reported from Iowa, North Dakota, and Ohio; I have not seen specimens from those states. It is adventive in northeastern California. Cirsium canescens hybridizes locally with C. scariosum and C. parryi. Further investigations may reveal that high-elevation forms of C. canescens from the mountains of Colorado are worthy of taxonomic recognition. These plants flower later than the low elevation forms of the Great Plains and occur in rather different ecologic conditions, but I have found no features that readily distinguish them. Populations of C. canescens have been particularly affected by the seedhead weevil Rhinocyllus conicus, introduced to North America to control weedy species of Carduus (S. M. Louda et al. 1997; Louda 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium pitcheri is endemic to beach and dune habitats around lakes Huron, Michigan, and Superior. It has been extirpated from portions of its former range at the southern end of Lake Michigan. It is threatened by foot traffic, off-road vehicular activity, and clearing and development of beachside habitats. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 122. | FNA vol. 19, p. 122. |
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium |
Sibling taxa | ||
Synonyms | C. nebraskense, C. plattense, C. nelsonii | Cnicus pitcheri |
Name authority | Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 420. (1841) | (Torrey ex Eaton) Torrey & A. Gray: Fl. N. Amer. 2: 456. (1843) |
Web links |