Flora of North America species comparison

several, erect to spreading, thinly arachnoid, soon glabrous;

branches ± fleshy, usually much branched proximally, spreading to ascending.

usually 1, erect, ± densely gray-tomentose with fine, non-septate trichomes;

branches 0 or few, usually above middle in distal 1/2, ascending.

blades ± elliptic, 30–75 × 10–20 cm, shallowly to deeply pinnatifid, lobes oblong to ovate, unlobed or with several prominent secondary lobes or large teeth, obtuse to acute, main spines 2–7 mm, abaxial gray arachnoid-tomentose, adaxial faces thinly arachnoid, glabrate;

basal often present at flowering, spiny winged-petiolate;

principal cauline sessile, bases clasping with broad, spiny-margined auricles, reduced distally, spinier than proximal;

distal much reduced, spines 7–20 mm.

blades oblong to elliptic or obovate, 10–25(–40) × 2–6(–12) cm, coarsely dentate or shallowly lobed to deeply pinnatifid, lobes well separated, triangular to linear or oblong, often revolute-margined, ascending to spreading, spinulose to spinose-dentate, main spines 2–3(–10) mm, faces gray-tomentose, more densely abaxially, sometimes glabrate adaxially;

basal usually present at flowering, winged-petiolate;

principal cauline progressively reduced distally, bases decurrent as spiny wings 1–5 cm, sometimes with expanded auricles;

distal cauline usually much reduced, less lobed.

0–7 cm.

0–10 cm.

ovoid to hemispheric or campanulate, 1.5–3 × 1.5–5 cm, sparsely to densely arachnoid, finely short-ciliate.

hemispheric to broadly campanulate, usually truncate or indented at base, 3–4 × 2.5–4 cm in first-formed heads, often smaller (1.5–2 cm) in later ones, loosely arachnoid on phyllary margins or glabrate.

dark reddish purple, 17–24 mm, tubes 8–11 mm, throats 3.5–6 mm, lobes 5–7 mm;

style tips 3–4 mm.

dull white or lavender-tinged, 20–35 mm, tubes 10–17 mm, throats 6–11 mm, lobes 4–9 mm;

style tips 5–8 mm.

in ca. 6 series, dark green or brown or with stramineous margins and a darker central zone, imbricate, linear-lanceolate (outer) to linear (inner), abaxial faces without glutinous ridge;

outer and mid bodies appressed, spiny-ciliate, apices long-spreading to ascending long-acuminate, spines straight, stout, 5–15 mm;

apices of inner straight or twisted, long, entire, flat or spine-tipped.

in 6–9 series, imbricate, ovate-lanceolate (outer) to linear-lanceolate (inner), abaxial faces with prominent glutinous ridge;

bodies of outer and middle appressed, acute, spines ascending to spreading, 2–4(–8) mm;

apices of inner expanded and flat, often twisted, scabrid-margined, and erose, spineless.

several–many, in congested corymbiform arrays.

1–10+, terminal on branches or in distal axils, in openly corymbiform to racemiform arrays.

dark brown, 4–5 mm, apical collars narrow;

pappi 15 mm.

light brown, 5–7 mm, sometimes with darker streaks, apical collar very narrow, lighter colored;

pappi 18–30 mm, usually noticeably shorter than corolla.

= 32.

= 34, 36.