Cirsium andersonii |
Cirsium eatonii |
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Anderson's thistle, rose thistle |
Eaton's thistle, mountaintop thistle, Peck's thistle, Steens Mountain thistle |
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Habit | Perennials (but often appearing biennial), (15–)40–70(–100) cm; rootstocks producing erect, taprooted caudices and rosettes. | Perennials, 10–150 cm; taprooted caudices. | ||||||||||||||||||||||||||||
Stems | usually 1, erect, subglabrous to puberulent and/or tomentose; branches 0–several, stiffly ascending. |
1–several, (fleshy), erect or ascending, simple to sparingly branched in distal 1/2, sometimes openly branched, glabrous to villous or tomentose with septate trichomes, sometimes ± glabrate; branches on distal stems 0–many, ascending. |
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Leaves | blades ± elliptic, 8–35 × 4–8 cm, divided about halfway to midveins, lobes spreading, triangular, coarsely dentate or with a few broad lobes, obtuse to acute, main spines 1–5 mm, abaxial faces green or gray, thinly tomentose, adaxial green and glabrous to sparingly pilose; basal often present at flowering, spiny winged-petiolate; main cauline reduced distally, bases clasping; distal much reduced, linear-oblong, usually less deeply lobed and often spinier than proximal. |
blades oblong, 10–30 × 1–5 cm, margins usually strongly undulate, unlobed and spiny-dentate or shallowly to deeply pinnatifid with 10–20 pairs of lobes, teeth or lobes closely spaced, often overlapping, lance-oblong to broadly triangular, deeply 3-lobed, ± spiny-dentate, main spines 2–12 mm, abaxial faces glabrous or villous with septate trichomes along midveins to densely arachnoid-tomentose, adaxial glabrous or villous with septate trichomes along midveins; basal often present at flowering, spiny winged-petiolate or sessile; principal cauline many, well distributed, proximally ± winged-petiolate, distally sessile, gradually reduced; distal not much reduced, often closely subtending heads. |
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Peduncles | 0–20 cm. |
0–14+ cm. |
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Involucres | broadly cylindric to narrowly campanulate, 3–5 × 2–4 cm, loosely arachnoid or ± glabrous, finely short-ciliate. |
green or suffused with dark purple, broadly ovoid to campanulate, 2–5 × 1.5–5 cm (appearing wider when pressed), loosely to densely villous or tomentose with septate trichomes and/or arachnoid-tomentose with finer, non-septate trichomes. |
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Corollas | red to reddish purple, 30–45 mm, tubes 10–20 mm, throats 10–16 mm, lobes 9–11 mm; style tips 3.5–5 mm. |
ochroleucous or yellow to lavender, pink, or purple, 15–35 mm, tubes 3.5–10 mm, throats 5–14 mm, lobes (linear), 4–12.5 mm; style tips 3–6 mm, conspicuously exserted beyond corolla lobes. |
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Phyllaries | in 6–8 series, imbricate, outer green, inner purple to red, linear-lanceolate (outer) to linear (inner), abaxial faces without glutinous ridge; outer and mid bodies short, appressed, entire or spinulose-ciliate, apices long-spreading to ascending, entire or spinulose-ciliate or rarely with expanded, fringed appendages, spines straight, weak, 1–3 mm; apices of inner red to purple, straight or rarely twisted, long, flat, entire. |
in 4–5 series, subequal, bases short-appressed, abaxial faces without or with very narrow glutinous ridge, apices usually stiffly ascending to spreading, linear-acicular, tapering to spines 7–35 mm; outer usually pinnately spiny, sometimes entire; apices of inner straight, plane or spine-tipped. |
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Heads | 1–6, borne singly or in corymbiform, racemiform, or spiciform arrays. |
1–many, erect or nodding, closely subtended by spiny-fringed bracts, usually sessile or short-pedunculate and crowded in subcapitate, spiciform, or racemiform (less commonly in openly branched) arrays. |
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Cypselae | brown, 6–7 mm, apical collars narrow; pappi 25–40 mm. |
dark brown, 5.5–7 mm, apical collars stramineous or not differentiated; pappi 12–25 mm. |
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2n | = 32, 64. |
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Cirsium andersonii |
Cirsium eatonii |
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Phenology | Flowering summer (Jul–Sep). | |||||||||||||||||||||||||||||
Habitat | Moist to dry soils, openings in montane woodlands, montaine coniferous forests, aspen groves | |||||||||||||||||||||||||||||
Elevation | 1100–2900 m (3600–9500 ft) | |||||||||||||||||||||||||||||
Distribution |
CA; ID; NV
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CO; ID; MT; NM; NV; OR; UT; WY; Rocky Mountains and high peaks of Great Basin desert region
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Discussion | Cirsium andersonii grows in the Cascade Range of northern California south through the Sierra Nevada of eastern California and western Nevada. It has been reported from the mountains of southwestern Idaho, but I have not seen specimens from there. Heads of Cirsium andersonii are actively visited by hummingbirds as well as a variety of insects (P. L. Barlow-Irick 2002). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 7 (7 in the flora). Cirsium eatonii is a polymorphic species widely distributed in a high elevation archipelago across the central Rocky Mountains and the Intermountain Region. During Pleistocene glacial episodes, the progenitors of this species complex undoubtedly occupied lower elevation sites and likely had more contiguous populations. Post-glacial isolation of these populations in allopatric high elevation sites has allowed them to differentiate to a greater or lesser extent. Prehistoric or recent introgressive hybridization with other thistle species probably has contributed to the diversification of the complex (R. J. Moore and C. Frankton 1965). Several of the races recognized here as varieties have been treated in the past as species (e.g., C. clokeyi, C. peckii). Their current geographic isolation and more or less distinctive features might support such recognition, but application of this approach across the complex would result in a proliferation of microspecies. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 145. | FNA vol. 19. | ||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium | ||||||||||||||||||||||||||||
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Synonyms | Cnicus andersonii | Cnicus eatonii | ||||||||||||||||||||||||||||
Name authority | (A. Gray) Petrak: Bot. Tiddsskr. 31: 68. (1911) | (A. Gray) B. L. Robinson: Rhodora 13: 240. (1911) | ||||||||||||||||||||||||||||
Web links |