Flora of North America species comparison

single, erect, villous with septate trichomes, sometimes ± glabrate, sometimes distally thinly tomentose;

branches few–many, ascending.

1–20+, erect or ascending, densely gray-tomentose with non-septate trichomes;

branches 0 or few, usually in distal 1/2, ascending.

blades oblanceolate to elliptic or ovate, 10–40 × 1–13 cm, margins flat, finely spiny-toothed and otherwise undivided to coarsely toothed or shallowly pinnatifid, lobes broadly triangular, main spines 1–5 mm, abaxial faces white-tomentose, adaxial faces green, glabrate to villous with septate trichomes;

basal usually absent at flowering, winged-petiolate, bases tapered;

principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping;

distal cauline well developed.

blades oblong to narrowly elliptic, 10–30 × 2–8 cm, strongly undulate, margins coarsely dentate or shallowly to deeply pinnatifid with 8–15 pairs of lobes 0.5–2 cm, often revolute, lobes ± triangular, closely spaced, spreading, spinose-dentate and cleft into 2–5 spine-tipped divisions, main spines 5–20 mm, yellowish, abaxial faces densely white-tomentose, adaxial thinly gray-tomentose;

basal usually present at flowering, winged-petiolate;

principal cauline sessile, progressively reduced distally, bases ± auriculate to long-decurrent as spiny wings;

distal cauline usually much reduced, less lobed.

0–5 cm (leafy-bracted).

0–4 cm.

ovoid to broadly cylindric or campanulate, (2–)2.5–3.5(–4) × (1.5–)2–3(–4) cm, thinly arachnoid.

ovoid to hemispheric or broadly campanulate, 2.5–4.5 × 2.5–4.5 cm in first-formed heads, often smaller in later ones, loosely arachnoid on phyllary margins or glabrate.

tips 4–6 mm.

pink to purple (white), 20–35 mm, tubes 10–16 mm, throats 5–12 mm, lobes 5–9 mm.

white or pale lavender to purple, pink, or red, 25–45 mm, tubes 8–25 mm, throats 6–17 mm, lobes 6–15 mm;

style tips 2–8 mm.

in 10–20 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with a narrow glutinous ridge (milky when fresh, dark when dry), outer and middle entire, bodies appressed, spines slender, abruptly spreading, 3–4 mm;

apices of inner phyllaries spreading, narrow, flattened, entire, spines spreading, slender, 3–4 mm;

apices of inner phyllaries spreading, narrow, flattened, ± dilated, ± erose or finely serrulate.

in 5–10 series, imbricate, ovate (outer) to linear-lanceolate (inner), margins entire, abaxial faces with narrow glutinous ridge;

outer and middle appressed, spines spreading, 3–12 mm;

apices of inner often flexuous, expanded and flat, scabrid-margined, sometimes erose, spineless.

1–many, in corymbiform or paniculiform arrays, (± elevated above principal cauline leaves), not subtended by ring of spiny bracts.

1–few, in leafy, ± corymbiform arrays.

tan to dark brown, 4–5.5 mm, apical collars stramineous, 0.5–1 mm;

pappi 12–24 mm.

light brown, sometimes with lighter or darker streaks, 6–9 mm, apical collars colored like the body, narrow;

pappi (white or tawny), 20–40 mm, usually noticeably shorter than corolla.

= 18.

= 30, 31, 32, 34.