Cirsium altissimum |
Cirsium andersonii |
|
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roadside thistle, tall thistle |
Anderson's thistle, rose thistle |
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Habit | Biennials or short-lived monocarpic perennials, (50–)100–300(–400) cm; taproots and often a cluster of coarse fibrous roots, roots without tuberlike enlargements. | Perennials (but often appearing biennial), (15–)40–70(–100) cm; rootstocks producing erect, taprooted caudices and rosettes. |
Stems | single, erect, villous with septate trichomes, sometimes ± glabrate, sometimes distally thinly tomentose; branches few–many, ascending. |
usually 1, erect, subglabrous to puberulent and/or tomentose; branches 0–several, stiffly ascending. |
Leaves | blades oblanceolate to elliptic or ovate, 10–40 × 1–13 cm, margins flat, finely spiny-toothed and otherwise undivided to coarsely toothed or shallowly pinnatifid, lobes broadly triangular, main spines 1–5 mm, abaxial faces white-tomentose, adaxial faces green, glabrate to villous with septate trichomes; basal usually absent at flowering, winged-petiolate, bases tapered; principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping; distal cauline well developed. |
blades ± elliptic, 8–35 × 4–8 cm, divided about halfway to midveins, lobes spreading, triangular, coarsely dentate or with a few broad lobes, obtuse to acute, main spines 1–5 mm, abaxial faces green or gray, thinly tomentose, adaxial green and glabrous to sparingly pilose; basal often present at flowering, spiny winged-petiolate; main cauline reduced distally, bases clasping; distal much reduced, linear-oblong, usually less deeply lobed and often spinier than proximal. |
Peduncles | 0–5 cm (leafy-bracted). |
0–20 cm. |
Involucres | ovoid to broadly cylindric or campanulate, (2–)2.5–3.5(–4) × (1.5–)2–3(–4) cm, thinly arachnoid. |
broadly cylindric to narrowly campanulate, 3–5 × 2–4 cm, loosely arachnoid or ± glabrous, finely short-ciliate. |
Style | tips 4–6 mm. |
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Corollas | pink to purple (white), 20–35 mm, tubes 10–16 mm, throats 5–12 mm, lobes 5–9 mm. |
red to reddish purple, 30–45 mm, tubes 10–20 mm, throats 10–16 mm, lobes 9–11 mm; style tips 3.5–5 mm. |
Phyllaries | in 10–20 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with a narrow glutinous ridge (milky when fresh, dark when dry), outer and middle entire, bodies appressed, spines slender, abruptly spreading, 3–4 mm; apices of inner phyllaries spreading, narrow, flattened, entire, spines spreading, slender, 3–4 mm; apices of inner phyllaries spreading, narrow, flattened, ± dilated, ± erose or finely serrulate. |
in 6–8 series, imbricate, outer green, inner purple to red, linear-lanceolate (outer) to linear (inner), abaxial faces without glutinous ridge; outer and mid bodies short, appressed, entire or spinulose-ciliate, apices long-spreading to ascending, entire or spinulose-ciliate or rarely with expanded, fringed appendages, spines straight, weak, 1–3 mm; apices of inner red to purple, straight or rarely twisted, long, flat, entire. |
Heads | 1–many, in corymbiform or paniculiform arrays, (± elevated above principal cauline leaves), not subtended by ring of spiny bracts. |
1–6, borne singly or in corymbiform, racemiform, or spiciform arrays. |
Cypselae | tan to dark brown, 4–5.5 mm, apical collars stramineous, 0.5–1 mm; pappi 12–24 mm. |
brown, 6–7 mm, apical collars narrow; pappi 25–40 mm. |
2n | = 18. |
= 32, 64. |
Cirsium altissimum |
Cirsium andersonii |
|
Phenology | Flowering summer–fall (Jun–Oct). | Flowering summer (Jul–Sep). |
Habitat | Prairies, woodlands, disturbed sites, often in damp soil | Moist to dry soils, openings in montane woodlands, montaine coniferous forests, aspen groves |
Elevation | 50–700 m (200–2300 ft) | 1100–2900 m (3600–9500 ft) |
Distribution |
AL; AR; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NY; OH; OK; PA; SC; SD; TN; TX; WI; WV
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CA; ID; NV
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Discussion | Plants of Cirsium altissimum ranging from southern Minnesota to Texas often have more deeply divided leaves than do populations in other portions of the species’ range. Some botanists (e.g., R. J. Moore and C. Frankton 1969; D. S. Correll and M. C. Johnston 1970) have treated those plants as C. iowense. Others (e.g., R. E. Brooks 1986; H. A. Gleason and A. Cronquist 1991; G. B. Ownbey and T. Morley 1991) have treated them as C. altissimum. Still others considered them to be derivatives of hybridization between C. altissimum and C. discolor (J. T. Kartesz and C. A. Meacham 1999) and treated them as C. ×iowense. Indeed the existence of these plants blurs the distinction between C. altissimum and C. discolor, and herbarium specimens are often difficult to assign. Natural hybrids between Cirsium altissimum and C. discolor are well documented (R. A. Davidson 1963; G. B. Ownbey and Hsi Y.-T. 1963; Ownbey 1964; S. Dabydeen 1997). Ownbey and Dabydeen both reported that apparent F1 hybrids between the two species have low seed set in comparison with the parental taxa. W. L. Bloom (1977) reported that the chromosomes of the two species differ by several rearareaments. Dabydeen reported a count of 2n = 19 with multiple meiotic irregularities for an apparent F1 hybrid. However, the presence of numerous individuals and populations seemingly intermediate between C. altissimum and C. discolor indicates that although F1 hybrids have low fertility, long-term processes may have stabilized hybrid derivatives of higher fertility. Ownbey and Hsi reported mitotic counts of 2n = 18 and 20 from a population that they treated as C. altissimum. In their discussion they noted that their plants represented “the segregate called C. iowense” and had been collected a short distance from that taxon’s type locality. R. J. Moore and C. Frankton (1969) reported a chromosome number of 2n = 18 for a plant from Texas that they considered to be C. iowense. Further investigation of morphologic variation, chromosome number, meiotic behavior, and fertility is needed of populations named as C. iowense to determine how those plants should be treated. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium andersonii grows in the Cascade Range of northern California south through the Sierra Nevada of eastern California and western Nevada. It has been reported from the mountains of southwestern Idaho, but I have not seen specimens from there. Heads of Cirsium andersonii are actively visited by hummingbirds as well as a variety of insects (P. L. Barlow-Irick 2002). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 111. | FNA vol. 19, p. 145. |
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium |
Sibling taxa | ||
Synonyms | Carduus altissimus, C. altissimum var. biltmoreanum, C. iowense | Cnicus andersonii |
Name authority | (Linnaeus) Sprengel: Syst. Veg. 3: 373. (1826) | (A. Gray) Petrak: Bot. Tiddsskr. 31: 68. (1911) |
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