Cicendia quadrangularis |
Gentianaceae |
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American cicendia, common microcalis, Oregon timwort, square cicendia, timwort |
gentian family |
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Habit | Herbs 1–9 cm. | Herbs [shrubs, trees], annual, biennial, or perennial, autotrophic, with green stems and leaves, or mycotrophic; when strongly mycotrophic, stems and leaves weakly chlorophyllous (only in Bartonia) or yellowish, whitish, purplish, or buff, or lacking chlorophyll (only in Voyria). | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stem | 1, branching at various levels in larger plants. |
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Leaves | basal often withered at flowering time, cauline spreading; basal blades spatulate-obovate to oblanceolate or elliptic-oblong, 3–15 × 1–3 mm; cauline blades narrowly elliptic-oblong to oblanceolate, 3.5–7 × 0.5–2 mm, base connate-sheathing 0.5 mm, apex obtuse to subacute. |
cauline, often also basal, opposite, whorled, or rarely alternate, sessile or petiolate, simple; stipules absent [rarely present as ocreae]; blade margins entire. |
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Inflorescences | 1–5-flowered; pedicels 3–60 mm. |
cymes (sometimes racemoid, spicoid, or capitate), thyrses, or verticillasters, or solitary flowers; flowers pedicellate or sessile. |
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Flowers | calyx hemispheric, 2–7 mm, much exceeding corolla tube in diam., outer lobes proximally rhombic, distal portion triangular, apex acute, inner lobes proximally oblong, distal portion with strongly invaginated flanges, abruptly narrowed into ovate-triangular, cucullate, acuminate apex; corolla 3–10 mm, lobes ovate-triangular to ovate-elliptic, 1–4 mm, apex rounded or abruptly acute. |
bisexual or occasionally some unisexual [all unisexual on some or all plants], homostylous [heterostylous], protandrous and outbreeding or less often homogamous and autogamous, radially [somewhat bilaterally] symmetric, 4–12(–14)-merous [rarely 3-, 6-, or 16-merous] except for carpels; perianth hypogynous, calyx and usually corolla persistent; calyx green or occasionally ± hyaline (absent in Obolaria), sepals connate or some [or all] nearly distinct, lobes imbricate in bud, often ± unequal, colleters often present adaxially near base; corolla petaloid, petals connate, lobes contorted in bud or rarely imbricate (Obolaria, Voyria), spurs present only in Halenia, 1 per petal; stamens epipetalous, isomerous and alternate with petals, all fertile [rarely some sterile], equal [unequal]; filaments free or connected by a corona; anthers 2-locular, dehiscing longitudinally [with terminal pores], remaining straight, recurving, or coiling helically or circinately, distinct or (only in some spp. |
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Fruits | capsular, dehiscence septicidal or rarely rupturing irregularly (Obolaria) [indehiscent capsules, berries]. |
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Seeds | few–very many, usually sessile; endosperm abundant and embryo small in autotrophic species, endosperm scant and embryo undifferentiated in completely mycotrophic species (Voyria). |
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Of | Gentiana) coherent; pistil 1, 2-carpellate; ovary 1[or 2]-locular; placentae 2, parietal [axile]; style present or absent, erect or initially deflexed to one side [declinate], uncleft, shallowly 2-cleft, or deeply cleft (Sabatia); stigmas 1 or 2, coiling only in Sabatia, decurrent on ovary (only in Lomatogonium, sometimes slightly so in Bartonia). |
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2n | = 26. |
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Cicendia quadrangularis |
Gentianaceae |
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Phenology | Flowering spring–early summer. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Grassy places, edges of vernal pools, other open habitats. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 0–2700 m. (0–8900 ft.) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; OR; South America [Introduced in Australia]
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nearly worldwide |
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Discussion | In the latter part of the life of the flower, the stamens of Cicendia quadrangularis arch over toward the stigmas, permitting self-pollination if outbreeding has not already occurred. The corollas close at night and in cloudy weather. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera ca. 100, species ca. 1800 (18 genera, 112 species in the flora). In the tribal classification by L. Struwe et al. (2002), genera 1–7 in this flora are in tribe Chironieae Dumortier, subtribe Chironiinae G. Don. Species of Chironieae generally lack nectaries, although Sabatia reportedly has indistinct nectaries at the base of the ovary. Genera 8–17 are in tribe Gentianeae Dumortier. Gentiana, in which the nectaries are on the gynophore, is in subtribe Gentianinae G. Don; the remaining genera of the Gentianeae in the flora area, all of which have epipetalous nectaries, are in subtribe Swertiinae Grisebach. Voyria constitutes the monogeneric tribe Voyrieae Gilg, in which the nectaries (when present) are on the ovary or the gynophore. Pedicel lengths given here refer to the true pedicels, between the most distal pair of bracts or bractlets and the calyx. In some genera, notably Centaurium, Sabatia, and Zeltnera, a flower terminating the ultimate branch of an inflorescence, directly subtended by bractlets, although sessile by this definition, may appear pedicellate. Corolla lengths as given are from the receptacle to the apices of the lobes (or plicae in Gentiana andrewsii). The Gentianaceae include many species esteemed in ornamental horticulture. In addition to those noted under the respective genera, the more important species in North American horticulture include Exacum affine Balfour f. ex Regel, Persian-violet, native to the island of Socotra, Yemen, which is widely grown as a florists’ pot plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 14. | FNA vol. 14. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Gentianaceae > Cicendia | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Gentiana quadrangularis, Microcala quadrangularis | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Dombey ex Lamarck) Grisebach: Gen. Sp. Gent., 157. (1838) | Jussieu | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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