Chylismia walkeri
Onagraceae subfam. onagroideae
Walker's evening-primrose, Walker's sun-cup
slender, unbranched or branched from base, 10–60 cm.
in basal rosette and/or cauline, often purple-dotted, 2–22 × 0.4–3.5 cm;
petiole 0.4–8 cm;
blade pinnately lobed, sometimes lateral lobes greatly reduced or absent and blade reduced to terminal lobe only, terminal lobe oblong or cordate to ovate, 1–5 × 0.5–3.2 cm, margins serrate, brown oil cells prominently lining veins abaxially.
stipules present or absent.
erect, elongating after anthesis.
opening at sunrise;
buds individually reflexed, with apical free tips less than 1 mm;
floral tube 0.5–1.5 mm, glabrous or sparsely villous inside;
sepals 1.5–5 mm;
petals bright yellow, fading pale orange or lavender, 1–6 mm;
stamens unequal, filaments of antisepalous stamens 1–3 mm, those of antipetalous ones 0.3–2 mm, anthers 0.5–2 mm, glabrous or sparsely ciliate;
style 1.5–6 mm, stigma surrounded by anthers at anthesis.
floral tube present or, rarely, absent;
sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens;
petals yellow, white, pink, red, rarely in combination.
spreading or ascending, oblong-cylindrical, 11–45 mm;
pedicel 5–30 mm.
0.6–1.2 mm.
Chylismia walkeri
Onagraceae subfam. onagroideae
Subspecies 2 (2 in the flora).
P. H. Raven (1962, 1969) determined this species to be self-incompatible and primarily autogamous.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 21, species 582 (16 genera, 246 species in the flora).
Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaves primarily cauline, rarely forming inconspicuous basal rosette, lateral lobes usually greatly reduced or absent; petals 1–3 mm; anthers 0.5–0.8 mm. | subsp. walkeri |
1. Leaves primarily basal, forming conspicuous rosette, cauline reduced or absent, lateral lobes usually well developed; petals 2.8–6 mm; anthers 1–2 mm. | subsp. tortilis |
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