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Walker's evening-primrose, Walker's sun-cup

Habit Herbs annual or short-lived perennial, villous, usually densely so proximally, less dense to glabrate distally, sometimes hairs somewhat appressed and shorter on leaves, also sometimes glandular puberulent on distal parts. Herbs perennial, base sometimes woody, glabrous or sparsely villous proximally.
Stems

slender, unbranched or branched from base, 10–60 cm.

with several divergent branches from base, 10–50 cm.

Leaves

in basal rosette and/or cauline, often purple-dotted, 2–22 × 0.4–3.5 cm;

petiole 0.4–8 cm;

blade pinnately lobed, sometimes lateral lobes greatly reduced or absent and blade reduced to terminal lobe only, terminal lobe oblong or cordate to ovate, 1–5 × 0.5–3.2 cm, margins serrate, brown oil cells prominently lining veins abaxially.

primarily in basal rosette and cauline, 3–20 × 0.7–2.5 cm;

petiole 0.5–4 cm;

blade pinnately or bipinnately lobed, terminal lobe ovate to elliptic, 0.4–2.5 × 0.3–1.5 cm, margins irregularly serrate, dark brown oil cells prominently lining veins abaxially.

Racemes

erect, elongating after anthesis.

erect, elongating after anthesis.

Flowers

opening at sunrise;

buds individually reflexed, with apical free tips less than 1 mm;

floral tube 0.5–1.5 mm, glabrous or sparsely villous inside;

sepals 1.5–5 mm;

petals bright yellow, fading pale orange or lavender, 1–6 mm;

stamens unequal, filaments of antisepalous stamens 1–3 mm, those of antipetalous ones 0.3–2 mm, anthers 0.5–2 mm, glabrous or sparsely ciliate;

style 1.5–6 mm, stigma surrounded by anthers at anthesis.

opening at sunrise;

buds individually reflexed, with apical free tips less than 1 mm;

floral tube 1.5–2 mm, glabrous inside;

sepals 2–5 mm;

petals bright yellow, with red dots near base, fading pale lavender, 2–6 mm;

stamens unequal, filaments of antisepalous stamens 1.5–3 mm, those of antipetalous ones 1–2 mm, anthers 1.2–2 mm, glabrous;

style 4–7 mm, stigma surrounded by anthers at anthesis.

Capsules

spreading or ascending, oblong-cylindrical, 11–45 mm;

pedicel 5–30 mm.

spreading to ascending, oblong-cylindrical, 8–20 mm;

pedicel 6–10 mm.

Seeds

0.6–1.2 mm.

0.6–1 mm.

Chylismia walkeri

Chylismia specicola

Distribution
sw United States
[WildflowerSearch map]
[BONAP county map]
Arizona
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

P. H. Raven (1962, 1969) determined this species to be self-incompatible and primarily autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

P. H. Raven (1962, 1969) thought that this species is most likely self-compatible but primarily outcrossing.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves primarily cauline, rarely forming inconspicuous basal rosette, lateral lobes usually greatly reduced or absent; petals 1–3 mm; anthers 0.5–0.8 mm.
subsp. walkeri
1. Leaves primarily basal, forming conspicuous rosette, cauline reduced or absent, lateral lobes usually well developed; petals 2.8–6 mm; anthers 1–2 mm.
subsp. tortilis
1. Leaf blades and styles glabrous; herbs ± woody at base.
subsp. specicola
1. Leaf blades and styles villous, at least at base; herbs not woody.
subsp. hesperia
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. walkeri
Subordinate taxa
C. walkeri subsp. tortilis, C. walkeri subsp. walkeri
C. specicola subsp. hesperia, C. specicola subsp. specicola
Synonyms Camissonia walkeri, Oenothera walkeri Oenothera specicola, Camissonia specicola
Name authority A. Nelson: Bot. Gaz. 56: 66. (1913) — (as Chylisma) (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 208. (2007)
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