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Walker's evening-primrose, Walker's sun-cup

redclay suncup

Habit Herbs annual or short-lived perennial, villous, usually densely so proximally, less dense to glabrate distally, sometimes hairs somewhat appressed and shorter on leaves, also sometimes glandular puberulent on distal parts. Herbs annual, sparsely to densely villous throughout or, sometimes, glabrate distally.
Stems

slender, unbranched or branched from base, 10–60 cm.

often intricately branched, 5–80 cm.

Leaves

in basal rosette and/or cauline, often purple-dotted, 2–22 × 0.4–3.5 cm;

petiole 0.4–8 cm;

blade pinnately lobed, sometimes lateral lobes greatly reduced or absent and blade reduced to terminal lobe only, terminal lobe oblong or cordate to ovate, 1–5 × 0.5–3.2 cm, margins serrate, brown oil cells prominently lining veins abaxially.

in poorly defined basal rosette and also cauline;

petiole 0.3–3.8 cm;

blade usually unlobed, very rarely pinnately lobed with few, small lateral lobes, ovate to elliptic, margins sparsely denticulate to subentire, pale or dark brown oil cells lining veins abaxially.

Racemes

erect, elongating after anthesis.

nodding, with intricate, filiform branches, elongating in fruit.

Flowers

opening at sunrise;

buds individually reflexed, with apical free tips less than 1 mm;

floral tube 0.5–1.5 mm, glabrous or sparsely villous inside;

sepals 1.5–5 mm;

petals bright yellow, fading pale orange or lavender, 1–6 mm;

stamens unequal, filaments of antisepalous stamens 1–3 mm, those of antipetalous ones 0.3–2 mm, anthers 0.5–2 mm, glabrous or sparsely ciliate;

style 1.5–6 mm, stigma surrounded by anthers at anthesis.

opening at sunrise;

buds without free tips;

floral tube 0.5–2 mm, glabrous or villous inside;

sepals 1.5–4 mm;

petals bright yellow, often with red dots near base, fading pale yellow or yellowish orange, 2–7 mm;

stamens unequal, filaments of antisepalous stamens 1.7–3.5 mm, those of antipetalous ones 1.2–2.5 mm, anthers 0.9–1.2 mm, glabrous;

style 4–9 mm, stigma exserted beyond anthers at anthesis.

Capsules

spreading or ascending, oblong-cylindrical, 11–45 mm;

pedicel 5–30 mm.

erect or ascending, clavate, 4–10 mm;

pedicel 4–20 mm.

Seeds

0.6–1.2 mm.

0.7–1.2 mm.

2n

= 14.

Chylismia walkeri

Chylismia parryi

Phenology Flowering May–Jun(–Sep).
Habitat Red clay and sand slopes weathered from red (freshwater-deposited) sandstone cliffs, with Juniperus or Larrea tridentata.
Elevation 800–1300 m. (2600–4300 ft.)
Distribution
sw United States
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

P. H. Raven (1962, 1969) determined this species to be self-incompatible and primarily autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chylismia parryi is known from northwestern Arizona (Coconino to Mohave counties) and southwestern Utah (Beaver to Washington counties), and is apparently disjunct to San Juan County, Utah. It is outcrossing and, perhaps, self-incompatible (P. H. Raven 1962, 1969). There are two morphological forms of this species. Raven (1962) noted that a later flowering form has narrower, smaller leaves, and less overall pubescence. It is not clear what these represent, but Raven (1962) made the combination Oenothera parryi forma tenuissima (M. E. Jones) P. H. Raven for it. He later (Raven 1969) noted that these plants did not seem to merit formal recognition, without any discussion.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves primarily cauline, rarely forming inconspicuous basal rosette, lateral lobes usually greatly reduced or absent; petals 1–3 mm; anthers 0.5–0.8 mm.
subsp. walkeri
1. Leaves primarily basal, forming conspicuous rosette, cauline reduced or absent, lateral lobes usually well developed; petals 2.8–6 mm; anthers 1–2 mm.
subsp. tortilis
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. scapoidea, C. specicola, C. walkeri
Subordinate taxa
C. walkeri subsp. tortilis, C. walkeri subsp. walkeri
Synonyms Camissonia walkeri, Oenothera walkeri Oenothera parryi, Camissonia parryi, C. tenuissima, O. scapoidea var. parryi, O. tenuissima
Name authority A. Nelson: Bot. Gaz. 56: 66. (1913) — (as Chylisma) (S. Watson) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma)
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