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Walker's evening-primrose, Walker's sun-cup

heart leaf primrose, heartleaf suncup

Habit Herbs annual or short-lived perennial, villous, usually densely so proximally, less dense to glabrate distally, sometimes hairs somewhat appressed and shorter on leaves, also sometimes glandular puberulent on distal parts. Herbs annual or perennial, villous and glandular puberulent.
Stems

slender, unbranched or branched from base, 10–60 cm.

usually well branched, forming bushy habit, 20–100 cm.

Leaves

in basal rosette and/or cauline, often purple-dotted, 2–22 × 0.4–3.5 cm;

petiole 0.4–8 cm;

blade pinnately lobed, sometimes lateral lobes greatly reduced or absent and blade reduced to terminal lobe only, terminal lobe oblong or cordate to ovate, 1–5 × 0.5–3.2 cm, margins serrate, brown oil cells prominently lining veins abaxially.

cauline, mostly toward base;

petiole (0.7–)2.5–7.5 cm;

blade cordate-ovate to -orbiculate, 2.5–7.5 × 2.3–5.5 cm, smaller distally, margins erose-dentate.

Racemes

erect, elongating after anthesis.

nodding, congested.

Flowers

opening at sunrise;

buds individually reflexed, with apical free tips less than 1 mm;

floral tube 0.5–1.5 mm, glabrous or sparsely villous inside;

sepals 1.5–5 mm;

petals bright yellow, fading pale orange or lavender, 1–6 mm;

stamens unequal, filaments of antisepalous stamens 1–3 mm, those of antipetalous ones 0.3–2 mm, anthers 0.5–2 mm, glabrous or sparsely ciliate;

style 1.5–6 mm, stigma surrounded by anthers at anthesis.

floral tube 4.5–14 mm, villous inside;

sepals 3–9 mm;

petals yellow, 3–12 mm;

filaments 1–3 mm, anthers 2–4 mm;

style 8–23 mm, stigma surrounded by or exserted just beyond anthers at anthesis.

Capsules

spreading or ascending, oblong-cylindrical, 11–45 mm;

pedicel 5–30 mm.

ascending, cylindrical, 20–55 mm;

pedicel 1–18 mm.

Seeds

0.6–1.2 mm.

0.5–0.7 mm.

Chylismia walkeri

Chylismia cardiophylla

Distribution
sw United States
[WildflowerSearch map]
[BONAP county map]
sw United States; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

P. H. Raven (1962, 1969) determined this species to be self-incompatible and primarily autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (2 in the flora).

P. H. Raven (1962, 1969) determined this species to be self-compatible, but primarily outcrossing. Subspecies cedrosensis (Greene) W. L. Wagner & Hoch, occurs in Baja California and adjacent Sonora, Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves primarily cauline, rarely forming inconspicuous basal rosette, lateral lobes usually greatly reduced or absent; petals 1–3 mm; anthers 0.5–0.8 mm.
subsp. walkeri
1. Leaves primarily basal, forming conspicuous rosette, cauline reduced or absent, lateral lobes usually well developed; petals 2.8–6 mm; anthers 1–2 mm.
subsp. tortilis
1. Herbs villous, sometimes also glandular puberulent; floral tubes 4.5–12 mm.
subsp. cardiophylla
1. Herbs glandular puberulent throughout, often also sparsely villous; floral tubes 9–14 mm.
subsp. robusta
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Lignothera
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola
C. arenaria, C. atwoodii, C. brevipes, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
Subordinate taxa
C. walkeri subsp. tortilis, C. walkeri subsp. walkeri
C. cardiophylla subsp. cardiophylla, C. cardiophylla subsp. robusta
Synonyms Camissonia walkeri, Oenothera walkeri Oenothera cardiophylla, Camissonia cardiophylla
Name authority A. Nelson: Bot. Gaz. 56: 66. (1913) — (as Chylisma) (Torrey) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma)
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