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Walker's sun cup

Habit Herbs (annual or perennial), [shrubs].
Leaves

primarily basal, forming conspicuous rosette, cauline reduced or absent;

blade usually pinnately lobed, rarely bipinnately, lateral lobes usually well developed.

alternate or basal;

stipules absent.

Flowers

floral tube 1–1.5 mm;

sepals 1.5–5 mm;

petals 2.8–6 mm;

anthers 1–2 mm, glabrous or sparsely ciliate;

style 4–6 mm.

usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;

stamens 2 times as many, or rarely as many, as sepals;

pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).

Fruit

a dry capsule, usually dehiscent, sometimes indehiscent.

Capsules

pedicel 5–30 mm.

Seeds

few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).

2n

= 14.

Chylismia walkeri subsp. tortilis

Onagraceae tribe Onagreae

Phenology Flowering Apr–Jun.
Habitat Colonial in rocky debris near cliffs, along ephemeral streams, on limestone.
Elevation 600–1900 m. (2000–6200 ft.)
Distribution
from FNA
CA; NV; UT
[BONAP county map]
North America; Mexico; Central America; South America; West Indies
Discussion

Subspecies tortilis is known from Millard and Washington counties, Utah, west to southern Elko and Mineral counties, Nevada, southwest to Inyo and northeastern San Bernardino counties, California, eastward to Clark County, Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 13, species 265 (12 genera, 199 species in the flora).

Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia > Chylismia walkeri Onagraceae > subfam. Onagroideae
Sibling taxa
C. walkeri subsp. walkeri
Subordinate taxa
Synonyms Oenothera scapoidea var. tortilis, Camissonia walkeri subsp. tortilis, C. walkeri var. tortilis, O. walkeri subsp. tortilis
Name authority (Jepson) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 208. (2007) Dumortier: Fl. Belg., 89. (1827)
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