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naked-stem beeblossom, Paiute suncup

Habit Herbs perennial, base sometimes woody, glabrous or sparsely villous proximally. Herbs annual, strigillose, villous, or glandular puberulent.
Stems

with several divergent branches from base, 10–50 cm.

usually unbranched, sometimes branched from base, 3–45 cm.

Leaves

primarily in basal rosette and cauline, 3–20 × 0.7–2.5 cm;

petiole 0.5–4 cm;

blade pinnately or bipinnately lobed, terminal lobe ovate to elliptic, 0.4–2.5 × 0.3–1.5 cm, margins irregularly serrate, dark brown oil cells prominently lining veins abaxially.

primarily in basal rosette, cauline poorly developed or absent, 1–18 × 0.5–3.5 cm;

petiole 0.5–6.5 cm;

blade pinnately lobed or lateral lobes greatly reduced or absent, sometimes mixed on same plant, terminal lobe narrowly ovate to ovate or elliptic, 1–6.5 × 0.5–3.3 cm, margins irregularly dentate to subentire, oil cells on abaxial surface inconspicuous or conspicuous, pale yellowish brown or dark brown.

Racemes

erect, elongating after anthesis.

nodding, elongating in fruit.

Flowers

opening at sunrise;

buds individually reflexed, with apical free tips less than 1 mm;

floral tube 1.5–2 mm, glabrous inside;

sepals 2–5 mm;

petals bright yellow, with red dots near base, fading pale lavender, 2–6 mm;

stamens unequal, filaments of antisepalous stamens 1.5–3 mm, those of antipetalous ones 1–2 mm, anthers 1.2–2 mm, glabrous;

style 4–7 mm, stigma surrounded by anthers at anthesis.

opening at sunrise;

buds with or without subapical free tips less than 1 mm;

floral tube 1–4 mm, sparsely villous or glabrous inside;

sepals 1.2–5 mm;

petals bright yellow, often with red dots near base, fading pale yellow or yellowish orange, 1.5–5.5(–8) mm;

stamens unequal, filaments of antisepalous stamens 1.2–6 mm, those of antipetalous ones 0.5–4 mm, anthers 1–2.5 mm, ciliate or glabrous;

style 3–11 mm, stigma surrounded by anthers at anthesis.

Capsules

spreading to ascending, oblong-cylindrical, 8–20 mm;

pedicel 6–10 mm.

ascending, clavate, 8–50 mm;

pedicel 4–20 mm.

Seeds

0.6–1 mm.

1–2 mm.

Chylismia specicola

Chylismia scapoidea

Distribution
Arizona
[BONAP county map]
w United States
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

P. H. Raven (1962, 1969) thought that this species is most likely self-compatible but primarily outcrossing.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

P. H. Raven (1962, 1969) determined this species to be self-compatible and primarily autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades and styles glabrous; herbs ± woody at base.
subsp. specicola
1. Leaf blades and styles villous, at least at base; herbs not woody.
subsp. hesperia
1. Oil cells lining veins of leaf blades abaxially usually pale yellowish brown, inconspicuous, rarely darker; leaf blades usually unlobed, rarely pinnately lobed with few, small lateral lobes, margins subentire.
→ 2
2. Capsules (10–)15–30 mm; petals 1.7–5 mm.
subsp. scapoidea
2. Capsules 25–50 mm; petals 1.5–2 mm.
subsp. macrocarpa
1. Oil cells lining veins of leaf blades abaxially dark brown, conspicuous; leaf blades pinnately lobed, lobes well developed or with few, small lateral lobes or unlobed, margins dentate.
→ 3
3. Petals 1.8–4 mm; lateral lobes of leaves often few or absent, sometimes well developed; capsules 8–20(–28) mm.
subsp. brachycarpa
3. Petals 4–5.5(–8) mm; lateral lobes of leaves usually well developed, sometimes poorly developed; capsules 16–38 mm.
subsp. utahensis
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. specicola, C. walkeri
Subordinate taxa
C. specicola subsp. hesperia, C. specicola subsp. specicola
C. scapoidea subsp. brachycarpa, C. scapoidea subsp. macrocarpa, C. scapoidea subsp. scapoidea, C. scapoidea subsp. utahensis
Synonyms Oenothera specicola, Camissonia specicola Oenothera scapoidea, Camissonia scapoidea, O. brevipes var. scapoidea
Name authority (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 208. (2007) (Torrey & A. Gray) Nuttall ex Raimann in H. G. A. Engler and K. Prantl: Nat. Pflanzenfam. 96[III,7]: 217. (1893)
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