Chylismia munzii |
Chylismia sect. Chylismia |
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Munz's evening primrose |
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Habit | Herbs annual, strigillose, often densely so. | Herbs usually annual, sometimes perennial, rarely biennial.Leaves basal and cauline, usually with well-developed basal rosette; blade usually pinnately or bipinnately lobed, sometimes with scattered, irregular lobes, sometimes lateral lobes greatly reduced or absent, terminal lobe elliptic, narrowly to broadly ovate to oblong, lanceolate, oblanceolate, cordate, or subcordate. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | several, 8–50 cm. |
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Leaves | primarily in basal rosette and also cauline, 1.5–20 × 0.5–3 cm; petiole 0.5–5 cm; blade pinnately lobed, terminal lobe ovate to narrowly ovate, 1.3–6 × 0.6–3 cm, margins denticulate, brownish oil cells lining veins abaxially. |
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Racemes | nodding, not congested, elongating in mature bud. |
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Flowers | opening at sunrise; buds with or without subapical free tips; floral tube orange-brown inside, 2–3 mm, villous inside; sepals 4–7 mm; petals bright yellow, with red dots near base, fading pale yellow or yellowish orange, 3–10 mm; stamens subequal, filaments 4–8 mm, anthers 3–6 mm, ciliate; style 8–18 mm, stigma exserted beyond anthers at anthesis. |
usually opening at sunrise, rarely at sunset; floral tube 0.4–9 mm; petals usually bright yellow, rarely white or cream, usually with red dots basally, or lavender to purple with white or yellow basally, sometimes with darker flecks near base, fading yellow, orange, reddish, or lavender; pollen shed singly. |
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Capsules | widely spreading, becoming sharply reflexed, clavate, 8–24 mm; pedicel 8–28 mm. |
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Seeds | 0.8–1.6 mm. |
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2n | = 14. |
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Chylismia munzii |
Chylismia sect. Chylismia |
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Phenology | Flowering Mar–Jun. | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Mesic slopes, washes. | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 600–1600 m. (2000–5200 ft.) | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; NV |
w United States; nw Mexico |
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Discussion | Chylismia munzii is known from middle elevations in the mountains at the north end, eastward from, and south of Death Valley, from Saline Valley and the Grapevine Mountains, Inyo County, California, and Yucca Flat, Nye County, Nevada, southward to the Kingston Range, San Bernardino County, California.P. H. Raven (1962, 1969) determined this species to be self-incompatible. It sometimes hybridizes with C. brevipes subsp. brevipes and C. claviformis subsp. aurantiaca. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 14 (14 in the flora). Section Chylismia consists of 10 diploid (2n = 14) species, and two that are partly polyploid (2n = 14, 28) [C. scapoidea subsp. scapoidea and C. walkeri subsp. walkeri (only one tetraploid population)]; no chromosome counts are available for the remaining two species, C. atwoodii and C. confertiflora (P. H. Raven 1962, 1969). Species of sect. Chylismia usually occur on sandy desert slopes, flats, and washes, often in sagebrush shrubland in the northern part of its range, or on rock slides or cliffs, mainly in the Mojave and northwestern Sonoran deserts, the Great Basin, and the lower elevations of the surrounding Sierra Nevada and Rocky Mountains. The limits of this range stretch from southeastern Oregon, central and southern Idaho, and central Wyoming, south through Nevada and Utah to eastern and southeastern California, northern Baja California and northwestern Sonora, Mexico, Arizona, northwestern New Mexico, and western Colorado. Several species are rare; C. confertiflora and C. specicola are known only from the Grand Canyon in northwestern Arizona, C. megalantha only from the vicinity of the type locality in Nye County, Nevada, and C. atwoodii also only from a narrow area around its type locality in Kane County, Utah. Others are widespread, especially the very diverse C. claviformis (11 subspp.), C. scapoidea (4 subspp.), C. walkeri (2 subspp.), and C. brevipes (3 subspp.). Chylismia scapoidea is the only species in the genus to occur east of the continental divide, both in Colorado on the upper Arkansas River in Fremont and Pueblo counties, and much more widely in Wyoming. Chylismia does not occur west of the Cascade-Sierra Nevada axis. Because R. A. Levin et al. (2004) included only C. claviformis in their analysis, they did not test the monophyly of sect. Chylismia; however, this section is both geographically distinct and morphologically set apart by the characteristic pinnate leaves (modified in some species, which have retained the entire apical lobe but do not have the smaller lateral lobes). Most species have bright yellow petals with red dots proximally and ultraviolet reflectance distally; some subspecies of C. claviformis have white petals; three species (C. atwoodii, C. heterochroma, and C. megalantha) have lavender or purple petals, often with lavender or purple flecks toward base, and white or yellow at the base and no reflectance, clearly a derived condition within the section (Raven 1962, 1969). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Oenothera munzii, Camissonia munzii | Camissonia section tetranthera, Oenothera section tetranthera | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 207. (2007) | unknown | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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