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froststem suncup

Habit Herbs annual or biennial, villous, at least proximally, glabrous or even glaucous distally, rarely glandular puberulent.
Stems

virgate with numerous divergent branches, 20–150 cm.

Leaves

primarily in well-developed basal rosette, cauline reduced or absent, 6–30 × 1.4–6.5 cm;

petiole 0.3–6 cm;

blade pinnately or bipinnately lobed, terminal lobe ovate to elliptic, 2.5–6.5 × 1.5–3 cm, margins irregularly serrate, dark brown oil cells prominently lining veins abaxially.

stipules present or absent.

Racemes

erect to nodding, elongating in bud.

Flowers

opening at sunrise;

buds individually reflexed, with apical or slightly subapical free tips less than 1 mm;

floral tube 1–3 mm, glabrous or villous inside proximally;

sepals 3–8 mm;

petals usually bright yellow, rarely cream, fading yellow to lavender, 4–9 mm;

stamens unequal, filaments of antisepalous stamens 2.5–4 mm, those of antipetalous ones 1.3–3 mm, anthers 2–4 mm, ciliate;

style 7–11 mm, stigma exserted beyond anthers at anthesis.

floral tube present or, rarely, absent;

sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens;

petals yellow, white, pink, red, rarely in combination.

Capsules

usually spreading, rarely slightly reflexed, oblong-cylindrical, 10–52 mm;

pedicel 7–20 mm.

Seeds

1–1.3 mm.

xI> = 7, 10, 11, 15, 18.

2n

= 14.

Chylismia multijuga

Onagraceae subfam. onagroideae

Phenology Flowering Mar–Jun(–Sep).
Habitat Forming colonies on rocky slopes and banks of eroded sedimentaries, on gypsum or limestone, on conglomerates, often with Juniperus and Pinus edulis, with Encelia farinosa and Larrea.
Elevation 300–1100 m. (1000–3600 ft.)
Distribution
from FNA
AZ; NV; UT
[WildflowerSearch map]
[BONAP county map]
North America; Mexico; Central America; South America; West Indies; Eurasia; Pacific Islands (New Zealand, Society Islands); Australia
Discussion

Chylismia multijuga is known from Washington County, Utah, and southern Lincoln County, Nevada, to northern Mohave County, Arizona. P. H. Raven (1962, 1969) determined this species to be self-incompatible. It hybridizes with C. brevipes subspp. brevipes and pallidula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 21, species 582 (16 genera, 246 species in the flora).

Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
Subordinate taxa
Synonyms Oenothera multijuga, Camissonia multijuga, Chylisma hirta, C. parviflora, C. venosa, O. brevipes var. multijuga, O. brevipes var. parviflora, O. multijuga var. parviflora, O. phlebophylla, O. watsonii
Name authority (S. Watson) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 41. (2007)
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