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froststem suncup

Habit Herbs annual or biennial, villous, at least proximally, glabrous or even glaucous distally, rarely glandular puberulent. Herbs perennial, base sometimes woody, glabrous or sparsely villous proximally.
Stems

virgate with numerous divergent branches, 20–150 cm.

with several divergent branches from base, 10–50 cm.

Leaves

primarily in well-developed basal rosette, cauline reduced or absent, 6–30 × 1.4–6.5 cm;

petiole 0.3–6 cm;

blade pinnately or bipinnately lobed, terminal lobe ovate to elliptic, 2.5–6.5 × 1.5–3 cm, margins irregularly serrate, dark brown oil cells prominently lining veins abaxially.

primarily in basal rosette and cauline, 3–20 × 0.7–2.5 cm;

petiole 0.5–4 cm;

blade pinnately or bipinnately lobed, terminal lobe ovate to elliptic, 0.4–2.5 × 0.3–1.5 cm, margins irregularly serrate, dark brown oil cells prominently lining veins abaxially.

Racemes

erect to nodding, elongating in bud.

erect, elongating after anthesis.

Flowers

opening at sunrise;

buds individually reflexed, with apical or slightly subapical free tips less than 1 mm;

floral tube 1–3 mm, glabrous or villous inside proximally;

sepals 3–8 mm;

petals usually bright yellow, rarely cream, fading yellow to lavender, 4–9 mm;

stamens unequal, filaments of antisepalous stamens 2.5–4 mm, those of antipetalous ones 1.3–3 mm, anthers 2–4 mm, ciliate;

style 7–11 mm, stigma exserted beyond anthers at anthesis.

opening at sunrise;

buds individually reflexed, with apical free tips less than 1 mm;

floral tube 1.5–2 mm, glabrous inside;

sepals 2–5 mm;

petals bright yellow, with red dots near base, fading pale lavender, 2–6 mm;

stamens unequal, filaments of antisepalous stamens 1.5–3 mm, those of antipetalous ones 1–2 mm, anthers 1.2–2 mm, glabrous;

style 4–7 mm, stigma surrounded by anthers at anthesis.

Capsules

usually spreading, rarely slightly reflexed, oblong-cylindrical, 10–52 mm;

pedicel 7–20 mm.

spreading to ascending, oblong-cylindrical, 8–20 mm;

pedicel 6–10 mm.

Seeds

1–1.3 mm.

0.6–1 mm.

2n

= 14.

Chylismia multijuga

Chylismia specicola

Phenology Flowering Mar–Jun(–Sep).
Habitat Forming colonies on rocky slopes and banks of eroded sedimentaries, on gypsum or limestone, on conglomerates, often with Juniperus and Pinus edulis, with Encelia farinosa and Larrea.
Elevation 300–1100 m. (1000–3600 ft.)
Distribution
from FNA
AZ; NV; UT
[WildflowerSearch map]
[BONAP county map]
Arizona
[BONAP county map]
Discussion

Chylismia multijuga is known from Washington County, Utah, and southern Lincoln County, Nevada, to northern Mohave County, Arizona. P. H. Raven (1962, 1969) determined this species to be self-incompatible. It hybridizes with C. brevipes subspp. brevipes and pallidula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

P. H. Raven (1962, 1969) thought that this species is most likely self-compatible but primarily outcrossing.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades and styles glabrous; herbs ± woody at base.
subsp. specicola
1. Leaf blades and styles villous, at least at base; herbs not woody.
subsp. hesperia
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. walkeri
Subordinate taxa
C. specicola subsp. hesperia, C. specicola subsp. specicola
Synonyms Oenothera multijuga, Camissonia multijuga, Chylisma hirta, C. parviflora, C. venosa, O. brevipes var. multijuga, O. brevipes var. parviflora, O. multijuga var. parviflora, O. phlebophylla, O. watsonii Oenothera specicola, Camissonia specicola
Name authority (S. Watson) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma) (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 208. (2007)
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