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Walker's evening-primrose, Walker's sun-cup

Habit Herbs annual, glandular pubescent throughout. Herbs annual or short-lived perennial, villous, usually densely so proximally, less dense to glabrate distally, sometimes hairs somewhat appressed and shorter on leaves, also sometimes glandular puberulent on distal parts.
Stems

several, 10–200 cm.

slender, unbranched or branched from base, 10–60 cm.

Leaves

in poorly defined basal rosette and cauline;

petiole 1.8–5.5 cm;

blade unlobed, broadly cordate to ovate, 2.4–8 × 7 cm, margins sinuate-dentate, yellowish oil cells prominently lining veins abaxially.

in basal rosette and/or cauline, often purple-dotted, 2–22 × 0.4–3.5 cm;

petiole 0.4–8 cm;

blade pinnately lobed, sometimes lateral lobes greatly reduced or absent and blade reduced to terminal lobe only, terminal lobe oblong or cordate to ovate, 1–5 × 0.5–3.2 cm, margins serrate, brown oil cells prominently lining veins abaxially.

Racemes

erect, elongating in flower.

erect, elongating after anthesis.

Flowers

opening at sunrise;

buds without free tips;

floral tube 4–9 mm, with matted, villous hairs inside;

sepals 4.5–9 mm;

petals pale to dark lavender, diffusely purplish-flecked near base, white at very base, fading darker lavender, 9–14 mm;

stamens unequal, filaments of antisepalous stamens 6–12 mm, of antipetalous ones 3.5–8 mm, anthers 2 mm, glabrous;

style 14–22.5 mm, stigma exserted beyond anthers at anthesis.

opening at sunrise;

buds individually reflexed, with apical free tips less than 1 mm;

floral tube 0.5–1.5 mm, glabrous or sparsely villous inside;

sepals 1.5–5 mm;

petals bright yellow, fading pale orange or lavender, 1–6 mm;

stamens unequal, filaments of antisepalous stamens 1–3 mm, those of antipetalous ones 0.3–2 mm, anthers 0.5–2 mm, glabrous or sparsely ciliate;

style 1.5–6 mm, stigma surrounded by anthers at anthesis.

Capsules

erect or ascending, clavate, 8–14 mm;

pedicel 2–3.5 mm.

spreading or ascending, oblong-cylindrical, 11–45 mm;

pedicel 5–30 mm.

Seeds

1–1.3 mm.

0.6–1.2 mm.

2n

= 14.

Chylismia megalantha

Chylismia walkeri

Phenology Flowering Jun–Oct.
Habitat Rubble derived from volcanic tuff, partly on moist soil along springs.
Elevation 1200–1400 m. (3900–4600 ft.)
Distribution
from FNA
NV
[BONAP county map]
sw United States
[WildflowerSearch map]
[BONAP county map]
Discussion

Chylismia megalantha is known from around Frenchman Drainage to French Peak and Skull Mountain in southern Nye County. P. H. Raven (1962, 1969) determined this species to be self-compatible, but outcrossing.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

P. H. Raven (1962, 1969) determined this species to be self-incompatible and primarily autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves primarily cauline, rarely forming inconspicuous basal rosette, lateral lobes usually greatly reduced or absent; petals 1–3 mm; anthers 0.5–0.8 mm.
subsp. walkeri
1. Leaves primarily basal, forming conspicuous rosette, cauline reduced or absent, lateral lobes usually well developed; petals 2.8–6 mm; anthers 1–2 mm.
subsp. tortilis
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola
Subordinate taxa
C. walkeri subsp. tortilis, C. walkeri subsp. walkeri
Synonyms Oenothera heterochromas. var. megalantha, Camissonia megalantha, O. megalantha Camissonia walkeri, Oenothera walkeri
Name authority (Munz) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 207. (2007) A. Nelson: Bot. Gaz. 56: 66. (1913) — (as Chylisma)
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