Chylismia heterochroma |
Onagraceae subfam. onagroideae |
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Shockley's evening-primrose |
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Habit | Herbs annual, glandular puberulent throughout, or glabrate and glaucous distally. | |
Stems | several, 10–100 cm. |
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Leaves | primarily in poorly defined basal rosette, cauline greatly reduced when present; petiole 0.4–8 cm; blade unlobed, ovate to cordate, 2–11.5 × 1.4–5 cm, margins sinuate-dentate, brown oil cells prominently lining veins abaxially. |
stipules present or absent. |
Racemes | erect, elongating in anthesis. |
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Flowers | opening at sunrise; buds without free tips; floral tube 2–5 mm, villous inside; sepals 1.5–3.5 mm; petals lavender, paler and often with flecks toward base, often yellow at very base, fading darker lavender, 2–6 mm; stamens unequal, filaments of antisepalous ones 1.8–3 mm, of antipetalous ones 1–2.5 mm, anthers 0.6–1 mm, glabrous or sparsely ciliate; style 4–7 mm, stigma surrounded by anthers at anthesis. |
floral tube present or, rarely, absent; sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens; petals yellow, white, pink, red, rarely in combination. |
Capsules | erect, clavate, 7–13 mm; pedicel 2–5 mm. |
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Seeds | 1–1.2 mm. |
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x |
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2n | = 14. |
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Chylismia heterochroma |
Onagraceae subfam. onagroideae |
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Phenology | Flowering May–Jun. | |
Habitat | Alluvial and rocky slopes. | |
Elevation | 600–2200 m. (2000–7200 ft.) | |
Distribution |
CA; NV
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North America; Mexico; Central America; South America; West Indies; Eurasia; Pacific Islands (New Zealand, Society Islands); Australia |
Discussion | Chylismia heterochroma is known from Churchill and Lander counties, Nevada, south to Lincoln and southern Nye counties, Nevada, to adjacent California (Mono Lake, Mono County, and central Inyo counties). P. H. Raven (1962, 1969) determined this species to be self-compatible and autogamous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 21, species 582 (16 genera, 246 species in the flora). Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia | Onagraceae |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Oenothera heterochromas., Camissonia heterochroma, C. heterochroma var. monoensis, O. heterochroma subsp. monoensis, O. heterochroma var. monoensis | |
Name authority | (S. Watson) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma) | W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 41. (2007) |
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