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Shockley's evening-primrose

Habit Herbs annual, glandular puberulent throughout, or glabrate and glaucous distally. Herbs perennial, base sometimes woody, glabrous or sparsely villous proximally.
Stems

several, 10–100 cm.

with several divergent branches from base, 10–50 cm.

Leaves

primarily in poorly defined basal rosette, cauline greatly reduced when present;

petiole 0.4–8 cm;

blade unlobed, ovate to cordate, 2–11.5 × 1.4–5 cm, margins sinuate-dentate, brown oil cells prominently lining veins abaxially.

primarily in basal rosette and cauline, 3–20 × 0.7–2.5 cm;

petiole 0.5–4 cm;

blade pinnately or bipinnately lobed, terminal lobe ovate to elliptic, 0.4–2.5 × 0.3–1.5 cm, margins irregularly serrate, dark brown oil cells prominently lining veins abaxially.

Racemes

erect, elongating in anthesis.

erect, elongating after anthesis.

Flowers

opening at sunrise;

buds without free tips;

floral tube 2–5 mm, villous inside;

sepals 1.5–3.5 mm;

petals lavender, paler and often with flecks toward base, often yellow at very base, fading darker lavender, 2–6 mm;

stamens unequal, filaments of antisepalous ones 1.8–3 mm, of antipetalous ones 1–2.5 mm, anthers 0.6–1 mm, glabrous or sparsely ciliate;

style 4–7 mm, stigma surrounded by anthers at anthesis.

opening at sunrise;

buds individually reflexed, with apical free tips less than 1 mm;

floral tube 1.5–2 mm, glabrous inside;

sepals 2–5 mm;

petals bright yellow, with red dots near base, fading pale lavender, 2–6 mm;

stamens unequal, filaments of antisepalous stamens 1.5–3 mm, those of antipetalous ones 1–2 mm, anthers 1.2–2 mm, glabrous;

style 4–7 mm, stigma surrounded by anthers at anthesis.

Capsules

erect, clavate, 7–13 mm;

pedicel 2–5 mm.

spreading to ascending, oblong-cylindrical, 8–20 mm;

pedicel 6–10 mm.

Seeds

1–1.2 mm.

0.6–1 mm.

2n

= 14.

Chylismia heterochroma

Chylismia specicola

Phenology Flowering May–Jun.
Habitat Alluvial and rocky slopes.
Elevation 600–2200 m. (2000–7200 ft.)
Distribution
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
Arizona
[BONAP county map]
Discussion

Chylismia heterochroma is known from Churchill and Lander counties, Nevada, south to Lincoln and southern Nye counties, Nevada, to adjacent California (Mono Lake, Mono County, and central Inyo counties). P. H. Raven (1962, 1969) determined this species to be self-compatible and autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

P. H. Raven (1962, 1969) thought that this species is most likely self-compatible but primarily outcrossing.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades and styles glabrous; herbs ± woody at base.
subsp. specicola
1. Leaf blades and styles villous, at least at base; herbs not woody.
subsp. hesperia
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. walkeri
Subordinate taxa
C. specicola subsp. hesperia, C. specicola subsp. specicola
Synonyms Oenothera heterochromas., Camissonia heterochroma, C. heterochroma var. monoensis, O. heterochroma subsp. monoensis, O. heterochroma var. monoensis Oenothera specicola, Camissonia specicola
Name authority (S. Watson) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma) (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 208. (2007)
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