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Shockley's evening-primrose

naked-stem beeblossom, Paiute suncup

Habit Herbs annual, glandular puberulent throughout, or glabrate and glaucous distally. Herbs annual, strigillose, villous, or glandular puberulent.
Stems

several, 10–100 cm.

usually unbranched, sometimes branched from base, 3–45 cm.

Leaves

primarily in poorly defined basal rosette, cauline greatly reduced when present;

petiole 0.4–8 cm;

blade unlobed, ovate to cordate, 2–11.5 × 1.4–5 cm, margins sinuate-dentate, brown oil cells prominently lining veins abaxially.

primarily in basal rosette, cauline poorly developed or absent, 1–18 × 0.5–3.5 cm;

petiole 0.5–6.5 cm;

blade pinnately lobed or lateral lobes greatly reduced or absent, sometimes mixed on same plant, terminal lobe narrowly ovate to ovate or elliptic, 1–6.5 × 0.5–3.3 cm, margins irregularly dentate to subentire, oil cells on abaxial surface inconspicuous or conspicuous, pale yellowish brown or dark brown.

Racemes

erect, elongating in anthesis.

nodding, elongating in fruit.

Flowers

opening at sunrise;

buds without free tips;

floral tube 2–5 mm, villous inside;

sepals 1.5–3.5 mm;

petals lavender, paler and often with flecks toward base, often yellow at very base, fading darker lavender, 2–6 mm;

stamens unequal, filaments of antisepalous ones 1.8–3 mm, of antipetalous ones 1–2.5 mm, anthers 0.6–1 mm, glabrous or sparsely ciliate;

style 4–7 mm, stigma surrounded by anthers at anthesis.

opening at sunrise;

buds with or without subapical free tips less than 1 mm;

floral tube 1–4 mm, sparsely villous or glabrous inside;

sepals 1.2–5 mm;

petals bright yellow, often with red dots near base, fading pale yellow or yellowish orange, 1.5–5.5(–8) mm;

stamens unequal, filaments of antisepalous stamens 1.2–6 mm, those of antipetalous ones 0.5–4 mm, anthers 1–2.5 mm, ciliate or glabrous;

style 3–11 mm, stigma surrounded by anthers at anthesis.

Capsules

erect, clavate, 7–13 mm;

pedicel 2–5 mm.

ascending, clavate, 8–50 mm;

pedicel 4–20 mm.

Seeds

1–1.2 mm.

1–2 mm.

2n

= 14.

Chylismia heterochroma

Chylismia scapoidea

Phenology Flowering May–Jun.
Habitat Alluvial and rocky slopes.
Elevation 600–2200 m. (2000–7200 ft.)
Distribution
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
w United States
[WildflowerSearch map]
[BONAP county map]
Discussion

Chylismia heterochroma is known from Churchill and Lander counties, Nevada, south to Lincoln and southern Nye counties, Nevada, to adjacent California (Mono Lake, Mono County, and central Inyo counties). P. H. Raven (1962, 1969) determined this species to be self-compatible and autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

P. H. Raven (1962, 1969) determined this species to be self-compatible and primarily autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Oil cells lining veins of leaf blades abaxially usually pale yellowish brown, inconspicuous, rarely darker; leaf blades usually unlobed, rarely pinnately lobed with few, small lateral lobes, margins subentire.
→ 2
2. Capsules (10–)15–30 mm; petals 1.7–5 mm.
subsp. scapoidea
2. Capsules 25–50 mm; petals 1.5–2 mm.
subsp. macrocarpa
1. Oil cells lining veins of leaf blades abaxially dark brown, conspicuous; leaf blades pinnately lobed, lobes well developed or with few, small lateral lobes or unlobed, margins dentate.
→ 3
3. Petals 1.8–4 mm; lateral lobes of leaves often few or absent, sometimes well developed; capsules 8–20(–28) mm.
subsp. brachycarpa
3. Petals 4–5.5(–8) mm; lateral lobes of leaves usually well developed, sometimes poorly developed; capsules 16–38 mm.
subsp. utahensis
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. specicola, C. walkeri
Subordinate taxa
C. scapoidea subsp. brachycarpa, C. scapoidea subsp. macrocarpa, C. scapoidea subsp. scapoidea, C. scapoidea subsp. utahensis
Synonyms Oenothera heterochromas., Camissonia heterochroma, C. heterochroma var. monoensis, O. heterochroma subsp. monoensis, O. heterochroma var. monoensis Oenothera scapoidea, Camissonia scapoidea, O. brevipes var. scapoidea
Name authority (S. Watson) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma) (Torrey & A. Gray) Nuttall ex Raimann in H. G. A. Engler and K. Prantl: Nat. Pflanzenfam. 96[III,7]: 217. (1893)
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