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Shockley's evening-primrose

redclay suncup

Habit Herbs annual, glandular puberulent throughout, or glabrate and glaucous distally. Herbs annual, sparsely to densely villous throughout or, sometimes, glabrate distally.
Stems

several, 10–100 cm.

often intricately branched, 5–80 cm.

Leaves

primarily in poorly defined basal rosette, cauline greatly reduced when present;

petiole 0.4–8 cm;

blade unlobed, ovate to cordate, 2–11.5 × 1.4–5 cm, margins sinuate-dentate, brown oil cells prominently lining veins abaxially.

in poorly defined basal rosette and also cauline;

petiole 0.3–3.8 cm;

blade usually unlobed, very rarely pinnately lobed with few, small lateral lobes, ovate to elliptic, margins sparsely denticulate to subentire, pale or dark brown oil cells lining veins abaxially.

Racemes

erect, elongating in anthesis.

nodding, with intricate, filiform branches, elongating in fruit.

Flowers

opening at sunrise;

buds without free tips;

floral tube 2–5 mm, villous inside;

sepals 1.5–3.5 mm;

petals lavender, paler and often with flecks toward base, often yellow at very base, fading darker lavender, 2–6 mm;

stamens unequal, filaments of antisepalous ones 1.8–3 mm, of antipetalous ones 1–2.5 mm, anthers 0.6–1 mm, glabrous or sparsely ciliate;

style 4–7 mm, stigma surrounded by anthers at anthesis.

opening at sunrise;

buds without free tips;

floral tube 0.5–2 mm, glabrous or villous inside;

sepals 1.5–4 mm;

petals bright yellow, often with red dots near base, fading pale yellow or yellowish orange, 2–7 mm;

stamens unequal, filaments of antisepalous stamens 1.7–3.5 mm, those of antipetalous ones 1.2–2.5 mm, anthers 0.9–1.2 mm, glabrous;

style 4–9 mm, stigma exserted beyond anthers at anthesis.

Capsules

erect, clavate, 7–13 mm;

pedicel 2–5 mm.

erect or ascending, clavate, 4–10 mm;

pedicel 4–20 mm.

Seeds

1–1.2 mm.

0.7–1.2 mm.

2n

= 14.

= 14.

Chylismia heterochroma

Chylismia parryi

Phenology Flowering May–Jun. Flowering May–Jun(–Sep).
Habitat Alluvial and rocky slopes. Red clay and sand slopes weathered from red (freshwater-deposited) sandstone cliffs, with Juniperus or Larrea tridentata.
Elevation 600–2200 m. (2000–7200 ft.) 800–1300 m. (2600–4300 ft.)
Distribution
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Chylismia heterochroma is known from Churchill and Lander counties, Nevada, south to Lincoln and southern Nye counties, Nevada, to adjacent California (Mono Lake, Mono County, and central Inyo counties). P. H. Raven (1962, 1969) determined this species to be self-compatible and autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chylismia parryi is known from northwestern Arizona (Coconino to Mohave counties) and southwestern Utah (Beaver to Washington counties), and is apparently disjunct to San Juan County, Utah. It is outcrossing and, perhaps, self-incompatible (P. H. Raven 1962, 1969). There are two morphological forms of this species. Raven (1962) noted that a later flowering form has narrower, smaller leaves, and less overall pubescence. It is not clear what these represent, but Raven (1962) made the combination Oenothera parryi forma tenuissima (M. E. Jones) P. H. Raven for it. He later (Raven 1969) noted that these plants did not seem to merit formal recognition, without any discussion.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. scapoidea, C. specicola, C. walkeri
Synonyms Oenothera heterochromas., Camissonia heterochroma, C. heterochroma var. monoensis, O. heterochroma subsp. monoensis, O. heterochroma var. monoensis Oenothera parryi, Camissonia parryi, C. tenuissima, O. scapoidea var. parryi, O. tenuissima
Name authority (S. Watson) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma) (S. Watson) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma)
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